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Genus Belonogaster de Saussure

Belonogaster de Saussure, 1854, Ét. Fam. Vesp. 2: 235, genus, replacement name for Raphigaster de Saussure, 1853.

Type species: Vespa juncea Fabricius, 1781, by subsequent designation of Bingham, 1897, Fauna Br. India, Hym. 1: 468.

Raphigaster de Saussure, 1853, Ét. Fam. Vesp. 2: 12, genus (7 species).  Non Raphigaster Laporte, 1832.

Belenogaster Walker, 1871, List Hym. Collected by J. K. Lord, Egypt and Arabia etc. (London): 39; Kirby, 1881, Proc. Zool. Soc. London: 649.  Incorrect spelling of Belonogaster de Saussure.

Lit.: Richards, 1982, Bull. Br. Mus. (Nat. Hist.) Entomol. 44: 31--114 (revision). -- Hensen and Blommers, 1987, Tijdschr. Entomol. 130: 11--31 (revision of Malagasy species). -- Gadagkar, 1991, in Ross and Matthews, Soc. Biol. Wasps: 149--190 (ethology).

 

Species list

 

abyssinica du Buysson

Raphigaster junceus var. de Saussure, 1853, Ét. Fam. Vesp. 2: pl. 2 fig. 2.

Belonogaster abyssinicus du Buysson, 1906, Bull. Soc. Entomol. France 1906: 190, female, male -- gAbyssinieh (lectotype female Paris); 1909, Ann. Soc. Entomol. France 78 (2): 214, 223, 263, pl. 4 fig. 4, pl. 5 fig. 6 (ethology). -- von Schulthess, 1912, Soc. Entomol. 27: 42 (compared to B. atratus n. sp.); 1913, Mitt. Zool. Mus. Berlin 6 (3): 338. -- Bequaert, 1918, Bull. Am. Mus. Nat. Hist. 39: 328 (cat.). -- Scott, 1933, Ann. Mag. Nat. Hist. 12: 120 (Ethiopia; Eritrea). -- Giordani Soika, 1939, Atti Mus. Civ. Stor. Nat. Trieste 14 (11): 170 (distr.); 1940, Atti Mus. Civ. Stor. Nat. Trieste 14 (18): 286 (distr.). -- Richards, 1982, Bull. Br. Mus. (Nat. Hist.) Entomol. 44: 40, 44--45 (key), 96, fig. 71 (abyssinica; designation of lectotype; distr.).

Distribution: Ethiopia; Eritrea; Djibouti.

 

acaulis Richards

Belonogaster acaulis Richards, 1982, Bull. Br. Mus. (Nat. Hist.) Entomol. 44: 34 (key), 61, fig. 18, female -- gSouth Africa: Natal National Parkh (London).

Distribution: South Africa.

 

adenensis adenensis Giordani Soika

Belonogaster abyssinicus adenensis Giordani Soika, 1957, Br. Mus. (Nat. Hist.), Exp. S. W. Arabia 1937--38, 1 (31): 484, female, male -- gWestern Aden Protectorate: Jebel Jihaf, 7100 fth (holotype female London); 1973, Boll. Mus. Civ. Stor. Nat. Venezia 24: 47.

Belonogaster adenensis adenensis; Richards, 1982, Bull. Br. Mus. (Nat. Hist.) Entomol. 44: 40, 46 (key), 99, fig. 77 (distr.); 1985 (1984), Fauna Saudi Arabia 6: 436, 437 (key), 438 (distr.).

Distribution: Yemen.

 

adenensis somaliensis Richards

Belonogaster adenensis somaliensis Richards, 1982, Bull. Br. Mus. (Nat. Hist.) Entomol. 44: 40, 46 (key), 100, female, male -- gSomali Republich (holotype female London); also from Ethiopia.

Distribution: Somalia; Ethiopia.

 

ambiko Hensen and Blommers

Belonogaster ambiko Hensen and Blommers, 1987, Tijdschr. Entomol. 130: 12 (key), 14, figs. 5--6, male -- gMadagascar; (Tamat.); Périnet; 950 m; 48‹16E, 18‹56Sh (Leiden). -- Kojima and van Achterberg, 1997, Zool. Meded., Leiden 71: 162 (holotype depository).

Distribution: Madagascar.

 

apicalis de Saussure

Belonogaster apicalis de Saussure, 1900, Abh. Senckenb. Naturforsch. Ges. 26 (2): 207 (key), 208, female -- gMadagascarh (holotype Genève). -- Dalla Torre, 1904, Genera Insectorum 19: 80 (cat.). -- du Buysson, 1909, Ann. Soc. Entomol. France 78 (2): 214, 230 (ethology). -- Bequaert, 1918, Bull. Am. Mus. Nat. Hist. 39: 329 (cat.). -- Richards, 1982, Bull. Br. Mus. (Nat. Hist.) Entomol. 44: 47(key), 105, figs. 85--86 (designation of "lectotype"; distr.). -- Hensen and Blommers, 1987, Tijdschr. Entomol. 130: 14 (key), 15. — Kojima, 2001a, Nat. Hist. Bull. Ibaraki Univ. (5): 11 (the specimen that Richards (1982) designated as the lectotype but not labeled is actually the unique holotype), 12 (Richards' males are different species; descriptionof male), figs. 13-21.

Distribution: Madagascar.

 

arabica Giordani Soika

Belonogaster grisea arabicus Giordani Soika, 1957, Br. Mus. (Nat. Hist.), Exp. S. W. Arabia 1937--38, 1 (31): 484, female, male -- gWestern Aden Protectorate: Dhala, 4800--5500 ft.h (holotype female London); also from other localities; 1973, Boll. Mus. Civ. Stor. Nat. Venezia 24: 47.

Belonogaster arabica; Richards, 1982, Bull. Br. Mus. (Nat. Hist.) Entomol. 44: 40, 45 (key), 96, figs. 72--73 (distr.).; 1985 (1984), Fauna Saudi Arabia 6: 436, 437 (key), 438 (distr.).

Distribution: Yemen; Saudi Arabia.

 

atrata von Schulthess

Belonogaster atratus von Schulthess, 1912, Soc. Entomol. 27: 41, female -- gWestafrika, Uelleburgh (lectotype Zürich); 1913, Mitt. Zool. Mus. Berlin 6 (3): 337, fig. 1 (female). -- Richards, 1982, Bull. Br. Mus. (Nat. Hist.) Entomol. 44: 34 (key), 59 (atrata; designation of lectotype).

Distribution: Cameroon.

 

aurata Richards

Belonogaster aurata Richards, 1982, Bull. Br. Mus. (Nat. Hist.) Entomol. 44: 34 (key), 59, fig. 14, female -- gNigeria: Lagosh (London).

Distribution: Nigeria.

 

barbata Richards

Belonogaster barbata Richards, 1982, Bull. Br. Mus. (Nat. Hist.) Entomol. 44: 39 (key), 92, fig. 66, female -- gCongo: Vokah (Paris).

Distribution: Congo.

 

betsileo Hensen and Blommers

Belonogaster brevipetiolata; Richards, 1982, Bull. Br. Mus. (Nat. Hist.) Entomol. 44: 101, fig. 80 (male) [partim].

Belonogaster betsileo Hensen and Blommers, 1987, Tijdschr. Entomol. 130: 12 (key), 15, figs. 7--8, male -- gBetsileo, Madagascarh (London).

Distribution: Madagascar.

 

bicolor de Saussure

Belonogaster bicolor de Saussure, 1900, Abh. Senckenb. Naturforsch. Ges. 26 (2): 207 (key), 208, female -- gMadagascarh (lectotype Genève). -- Dalla Torre, 1904, Genera Insectorum 19: 80 (cat.). -- Richards, 1982, Bull. Br. Mus. (Nat. Hist.) Entomol. 44: 47 (key), 104 (designation of lectotype; distr.). -- Hensen and Blommers, 1987, Tijdschr. Entomol. 130: 12 (key), 16, fig. 10 (syn.: malagassus de Saussure; distr.). — Kojima, 2001a, Nat. Hist. Bull. Ibaraki Univ. (5): 12 (bicolor and malagassa are different secies), 13 (key), figs. 23, 25-26, 28..

Belonogaster prasinus var. bicolor; du Buysson, 1909, Ann. Soc. Entomol. France 78 (2): 229. -- Bequaert, 1918, Bull. Am. Mus. Nat. Hist. 39: 336 (cat.).

Distribution: Madagascar.

 

bimaculata Richards

Belonogaster bimaculata Richards, 1982, Bull. Br. Mus. (Nat. Hist.) Entomol. 44: 37, 45 (key), 78, figs. 49--50, female, male -- gZambia: Abercornh (holotype female London); also from 6 other localities in Zambia.

Distribution: Zambia.

 

brachystoma Kohl

Belonogaster brachystomus Kohl, 1894 (1893), Ann. Naturhist. Hofmus. Wien 9: 322 (key), 326, 334, pl. 15 fig. 78, pl. 17 fig. 136, female, male -- gOstafrika Delagoa-Bayh (lectotype male Wien). -- Dalla Torre, 1894, Cat. Hym. 9: 114 (brachystoma; cat.); 1904, Genera Insectorum 19: 80 (brachystoma; cat.). -- du Buysson, 1909, Ann. Soc. Entomol. France 78 (2): 217, 219, 252, 265. -- von Schulthess, 1914, Dtsch. Entomol. Z. 1914: 290 (compared to B. rothkirchi n. sp.). -- Bequaert, 1918, Bull. Am. Mus. Nat. Hist. 39: 329 (cat.). -- Giordani Soika, 1961, S. Afr. Anim. Life 8: 450 (distr.). -- Richards, 1982, Bull. Br. Mus. (Nat. Hist.) Entomol. 44: 39, 41 (key), 68, figs. 33--34 (brachystoma; distr.).

Distribution: Zambia; Zimbabwe; Tanzania (including Zanzibar); Malawi; Mozambique; South Africa (Kwazulu-Natal, Transvaal).

 

brevipetiolata de Saussure

Belonogaster brevipetiolatus de Saussure, 1890, in Grandidier, Hist. Madagascar 20, Hym. 1: 89 (key), 98, pl. 4 fig. 1, female, male -- gMadagascar. -- Fianarantsoa (pays des Betsileo.) et environs dfAndgrangoloaka (à la limite est de la province dfImerina) ... Còte sud ... Nosibéh (Genève). -- Kohl, 1894 (1893), Ann. Naturhist. Hofmus. Wien 9: 321, 323 (key), 334, pl. 15 fig. 83, pl. 16 figs. 95, 99, 120. -- Gribodo, 1894, in Emery et al., Mem. R. Accad. Sci. Bologna (5) 4: 123. -- Dalla Torre, 1894, Cat. Hym. 9: 114 (brevipetiolata; cat.). -- de Saussure, 1900, Abh. Senckenb. Naturforsch. Ges. 26 (2): 207 (key). -- Dalla Torre, 1904, Genera Insectorum 19: 80 (brevipetiolata; cat.). -- du Buysson, 1909, Ann. Soc. Entomol. France 78 (2): 214, 227, 263, pl. 6 fig. 1 (ethology). -- Bequaert, 1918, Bull. Am. Mus. Nat. Hist. 39: 329 (cat.). -- van der Vecht, 1968, Proc. K. Ned. Akad. Wet. Ser. C Biol. Med. Sci. 71: fig. 4d (brevipetiolata; morphology). -- Richards, 1982, Bull. Br. Mus. (Nat. Hist.) Entomol. 44: 33, 46 (key), 101, fig. 79 (brevipetiolata; nest; distr.) [partim, not male]. -- Hensen and Blommers, 1987, Tijdschr. Entomol. 130: 12 (key), 16, figs. 11, 12 (brevipetiolata; compared to B. erythrocephala n. sp.; distr.). — Kojima, 2001a, Nat. Hist. Bull. Ibaraki Univ. (5): 10 (notes on lectotype and paralectotypes).

Distribution: Madagascar.

 

brevitarsus Richards

Belonogaster brevitarsus Richards, 1982, Bull. Br. Mus. (Nat. Hist.) Entomol. 44: 35, 39, 42 (key), 92, figs. 67, 68, male, female -- gUganda: Mabira forest, Chagwe, 3500--3800 ft, [1070--1160 m]h (holotype male London); also from two other localities in Uganda; and Kenya; Zaire; Rwanda.

Distribution: Uganda; Kenya; Rwanda; Zaire.

 

brunnea brunnea Ritsema

Belonogaster brunneus Ritsema, 1874, Tijdschr. Entomol. 17: 177, 202, female -- gNeder-Guineah [labeled gCongoh (= Congo River)] (unique Leiden). -- Kohl, 1894 (1893), Ann. Naturhist. Hofmus. Wien 9: 334. -- Dalla Torre, 1894, Cat. Hym. 9: 114 (brunea [!]; cat. [erroneously recorded from gAustr., New Guineah]). -- Gribodo, 1895, Mem. R. Accad. Sci. Bologna (5) 5: 335. -- Dalla Torre, 1904, Genera Insectorum 19: 80 (brunea [!]; cat.; recorded from gUnter-Guineah). -- du Buysson, 1909, Ann Soc. Entomol. France 78 (2): 217, 219, 245, 264, pl. 6 fig. 2 (male; ethology); 1914, in Voy. Alluaud et Jeannel Afr. Or., Rés. Sci., Ins. Hym. 3, Vesp.: 162. -- Schulz, 1906, Spolia Hym.: 226. -- Bequaert, 1918, Bull. Am. Mus. Nat. Hist. 39: 234, 329 (cat.; syn.: B. distinguendus Kohl). -- Salt and Bequaert, 1929, Psyche 36: 262 (distr.; record of stylopized specimen). -- Bequaert, 1938, Explor. Parcs Nation. Congo Belge Miss. G. F. de Witte (1933--1935) 11: 8, 9, 10 (distr.). -- Benoit, 1956, Ann. Mus. R. Congo Belge 8 (Sci. Zool.) 51: 552 (distr.). -- Kojima and van Achterberg, 1997, Zool. Meded., Leiden 71: 162 (holotype depository).

Belonogaster distinguendus Kohl, 1894 (1893), Ann. Naturhist. Hofmus. Wien 9: 322, 323 (key), 328, 335, pl. 15 figs. 79--81, 94, pl. 16 fig. 98, pl. 17 fig. 141, female, male -- gWestafrika: Chutes de Samlia, Riv. N. Gamio, Mocquereys Grand Bana ...h (lectotype male Wien). -- Dalla Torre, 1894, Cat. Hym. 9: 114 (distinguenda; cat.); 1904, Genera Insectorum 19: 80 (distinguenda; cat.). -- Schulz, 1905, Hym. Studien: 15. -- Bequaert, 1918, Bull. Am. Mus. Nat. Hist. 39: 329 (syn. of B. brunneus Ritsema). -- Richards, 1982, Bull. Br. Mus. (Nat. Hist.) Entomol. 44: 64 (designation of lectotype; marked as gSyn. n.h).

Belonogaster brunnea brunnea; Richards, 1982, Bull. Br. Mus. (Nat. Hist.) Entomol. 44: 38, 41--43 (key), 64, figs. 23--25 (distr.).

Distribution: Sierra Leone; Liberia; Ghana; Nigeria; Bioko; Gabon; Congo; Zaire; Rwanda; Burundi.

 

brunnea nigriclava Richards

Belonogaster brunnea nigriclava Richards, 1982, Bull. Br. Mus. (Nat. Hist.) Entomol. 44: 38, 43 (key), 66, figs. 26--27, male, female -- gUganda: Mt Kokanjero, SW. Mt Elgon, 6400 ft [1950m]h (holotype male London); also from 12 other localities in Uganda; and Kenya; Zambia; Malawi; Tanzania; Zaire. — Kojima, 2001a, Nat. Hist. Bull. Ibaraki Univ. (5): 20.

Distribution: Uganda; Kenya; Zaire; Tanzania; Zambia; Malawi.

 

brunnescens Richards

Belonogaster brunnescens Richards, 1982, Bull. Br. Mus. (Nat. Hist.) Entomol. 44: 38, 45 (key), 70, male, female -- gZimbabwe: Mashonalandh (holotype male London); also from Ethiopia; Gabon; Congo; Zaire; Zimbabwe; Zambia; Kenya; Uganda; Mozambique; Malawi; Tanzania (Zanzibar); South Africa (Transvaal).

Distribution: Ethiopia; Kenya; Uganda; Tanzania (including Zanzibar); Congo; Zaire; Gabon; Malawi; Zambia; Zimbabwe; Mozambique; South Africa (Transvaal).

 

clypeata clypeata Kohl

Belonogaster clypeatus Kohl, 1894 (1893), Ann. Naturhist. Hofmus. Wien 9: 321 (key), 328, 335, pl. 15 figs. 88, 91, pl. 17 figs. 124, 144, 149, female, male -- gDeutsch-Mozambiqueh (lectotype male Wien); also from Madagascar [error]. -- Dalla Torre, 1894, Cat. Hym. 9: 114 (clypeata; cat.). -- Stadelmann, 1898, Deutsch Ost-Afrika 4, Hym.: 35. -- von Schulthess, 1899, Bull. Soc. Vaud. Sci. Nat. (4) 35: 269 (distr.). -- Dalla Torre, 1904, Genera Insectorum 19: 80 (clypeata; cat.). -- du Buysson, 1909, Ann. Soc. Entomol. France 78 (2): 217, 219, 244, 264, pl. 2 figs. 9, 10, pl. 5 figs. 1, 2 (ethology); 1914, in Voy. Alluaud et Jeannel Afr. Or., Rés. Sci., Ins. Hym. 3, Vesp.: 162. -- Bequaert, 1918, Bull. Am. Mus. Nat. Hist. 39: 329 (cat.). -- Schouteden, 1919, Rev. Zool. Afr. 6: 185 (distr.). -- Giordani Soika, 1935, Mem. Estud. Mus. Zool. Univ. Coimbra (1) 82: 14 (distr.). -- Bequaert, 1938, Explor. Parcs Nation. Congo Belge Miss. G. F. de Witte (1933--1935) 11: 9 (distr.). -- Guiglia, 1940, Boll. Lab. Zool. Gen. Agrar. Fac. Agrar. Portici 31: 278 (distr.). -- Richards, 1982, Bull. Br. Mus. (Nat. Hist.) Entomol. 44: 67 (designation of lectotype).

Belonogaster clypeata clypeata; Richards, 1982, Bull. Br. Mus. (Nat. Hist.) Entomol. 44: 34, 41 (key), 67, figs. 28--31 (distr.).

Distribution: Uganda; Tanzania (including Zanzibar); Zaire; Malawi; Zambia; Zimbabwe; Mozambique; South Africa (Transvaal, Kwazulu-Natal, Western Cape).

 

clypeata fuscata Richards

Belonogaster clypeata fuscata Richards, 1982, Bull. Br. Mus. (Nat. Hist.) Entomol. 44: 34, 41 (key), 67, female, male -- gUganda: Ankole‑Toro border, east of Lake George, 4500 ft [1370 m]h (holotype female London); also from five other localities in Uganda; and Zambia; Angola; Zaire.

Distribution: Uganda; Zaire; Zambia; Angola.

 

confusa Kojima

Belonogaster pictus de Saussure, 1900, Abh. Senckenb. Naturforsch. Ges. 26 (2): 207 (key) [Junior primary homonym of Belonogaster pictus Kohl, 1894]. — Kojima, 2001a, Nat. Hist. Bull. Ibaraki Univ. (5): 16 (designation of lectotype).

Belonogaster hildebrandti var.; de Saussure, 1900, Abh. Senckenb. Naturforsch. Ges. 26 (2): 210

Belonogaster confusa Kojima, 2001a: 16, figs. 54-60, 19 (key). Replacement name for Belonogaster pictus de Saussure, 1900.

Distribution: Madagascar.

 

dayi Hensen and Blommers

Belonogaster dayi Hensen and Blommers, 1987, Tijdschr. Entomol. 130: 12 (key), 16, figs. 9, 36, 37, 42, male, female -- gMadagascar: Tamat.; Périneth (holotype female London); also from 3 other localities in Madagascar; nest is also noted.

Distribution: Madagascar.

 

discifera Hensen and Blommers

Belonogaster discifera Hensen and Blommers, 1987, Tijdschr. Entomol. 130: 14 (key), 18, figs. 13, 14, male, female -- gRép. Malgache, Manjakandriana, Mandraka, 1300 mh (holotype male Amsterdam); also from other locality in Madagascar. -- Wenzel, 1991, in Ross and Matthews, Soc. Biol. Wasps: 506 (nest). -- Kojima and van Achterberg, 1997, Zool. Meded., Leiden 71: 166 (holotype depository).

Distribution: Madagascar.

 

dubia Kohl

Belonogaster dubius Kohl, 1894 (1893), Ann. Naturhist. Hofmus. Wien 9: 322, 323 (key), 329, 335, pl. 15 figs. 79, 80, 94, pl. 16 fig. 100, pl. 18 fig. 145, female, male -- gOstafrika: Dar es Salaamh (lectotype male Wien). -- Dalla Torre, 1894, Cat. Hym. 9: 114 (dubia; cat.). -- Stadelmann, 1898, Deutsch Ost-Afrika 4, Hym.: 35 -- Dalla Torre, 1904, Genera Insectorum 19: 80 (dubia; cat.). -- du Buysson, 1909, Ann. Soc. Entomol. France 78 (2): 215, 219, 235, 263 (ethology). -- Roubaud, 1910, C. R. Acad. Sci. Paris 151: 553--556 (ethology). -- von Schulthess, 1913, Mitt. Zool. Mus. Berlin 6 (3): 338; 1913, Ark. Zool. 8 (17): 14 (syn.: B. massaicus Cameron). -- du Buysson, 1914, in Voy. Alluaud et Jeannel Afr. Or., Rés. Sci., Ins. Hym. 3, Vesp.: 161. -- Bequaert, 1918, Bull. Am. Mus. Nat. Hist. 39: 235 (key), 236 (ethology), 330 (cat.). -- Schouteden, 1919, Rev. Zool. Afr. 6: 184 (distr.). -- von Schulthess; 1929, Rev. Zool. Bot. Afr. 17 (2): 187 (distr.). -- Salt and Bequaert, 1929, Psyche 36: 262 (record of stylopized specimen). -- Bequaert, 1938, Explor. Parcs Nation. Congo Belge Miss. G. F. de Witte (1933--1935) 11: 8, 9, 10 (distr.). -- Guiglia, 1940, Boll. Lab. Zool. Gen. Agrar. Fac. Agrar. Portici 31: 279 (distr.). -- Marino Piccioli, 1968, Monit. Zool. Ital. (n. ser.) 2, Suppl.: 203 (ethology). -- Richards, 1982, Bull. Br. Mus. (Nat. Hist.) Entomol. 44: 35, 38, 43 (key), 53, figs. 10, 11 (dubia; designation of lectotype; syn.: B. occidentalis Tullgren, B. massaicus Cameron; distr.). -- Kojima and Keeping, 1988, Kontyû, 56: 817 (dubia; larva).

Belonogaster saevus var. de Saussure, 1890, in Grandidier, Hist. Madagascar 20, Hym. 1: 92, in footnote (female).

Belenogaster [!] occidentalis Tullgren, 1904, Ark. Zool. 1: 455, pl. 25, fig. 14, female, male -- [Cameroon] gItokih (lectotype male Stockholm); also from Bonge, Ekundu; Tanzania: Dar-es-Salaam. -- Bequaert, 1918, Bull. Am. Mus. Nat. Hist. 39: 336 (syn. of B. saevus de Saussure). -- Richards, 1982, Bull. Br. Mus. (Nat. Hist.) Entomol. 44: 54 (Belonogaster; designation of lectotype).

Belonogaster massaicus Cameron, 1910, in Sjöstedt, Wiss. Ergebn. Schwed. Zool. Exp. Kilimandjaro 2 (8), Hym. 6: 171, female -- gKilimandjaro: Kibonoto 1000--1200 mh [Tanzania] (lectotype Stockholm). -- von Schulthess, 1913, Ark. Zool. 8 (17): 14 (syn. of B. dubius). -- Richards, 1982, Bull. Br. Mus. (Nat. Hist.) Entomol. 44: 54 (designation of lectotype; marked as gSyn. n.h).

Distribution: Senegal; Gambia; Guinea; Liberia; Côte dfIvoire; Ghana; Niger; Nigeria; Bioko; Congo; Zaire; Uganda; Rwanda; Kenya; Tanzania; Angola; Zambia; Malawi; Zimbabwe; Mozambique; South Africa (Transvaal, Kwazulu-Natal).

 

erythrocephala Hensen and Blommers

Belonogaster brevipetiolata; de Saussure, 1890, in Grandidier, Hist. Madagascar 20, Hym. 1: 98, pl. 4 fig. 1 [partim]. -- Richards, 1982, Bull. Br. Mus. (Nat. Hist.) Entomol. 44: 101, fig. 79, female [partim].

Belonogaster erythrocephala Hensen and Blommers, 1987, Tijdschr. Entomol. 130: 12 (key), 19, figs. 15, 16, male, female -- gRép. Malgache, Nosy Be, Dzamandzarh (holotype male Amsterdam); also from two other localities in Madagascar. -- Kojima and van Achterberg, 1997, Zool. Meded., Leiden 71: 167 (holotype depository).

Distribution: Madagascar.

 

eumenoides de Saussure

Belonogaster eumenoides de Saussure, 1890, in Grandidier, Hist. Madagascar 20, Hym. 1: 89 (key), 94, pl. 17 fig. 12, female -- gMadagascar ... aux environs dfAndrangoloakah (lectotype Genève); also from another locality. -- Dalla Torre, 1894, Cat. Hym. 9: 114 (cat.). -- Kohl, 1894 (1893), Ann. Naturhist. Hofmus. Wien 9: 335. -- de Saussure, 1900, Abh. Senckenb. Naturforsch. Ges. 26 (2): 207 (key), 209. -- Dalla Torre, 1904, Genera Insectorum 19: 80 (cat.). -- du Buysson, 1909, Ann Soc. Entomol. France 78 (2): 215, 234, 263, pl. 4 fig. 6 (male; ethology). -- Bequaert, 1918, Bull. Am. Mus. Nat. Hist. 39: 330 (syn.: pomicolor de Saussure, ornatus de Saussure; cat.). -- Richards, 1982, Bull. Br. Mus. (Nat. Hist.) Entomol. 44: 47 (key), 110, figs. 91, 92 (designation of lectotype; ?syn.: malagassus de Saussure; distr.) [partim]. -- Hensen and Blommers, 1987, Tijdschr. Entomol. 130: 14 (key), 20, figs. 2, 17--19, 43 (female, male; nest; distr.). — Kojima, 2001a, Nat. Hist. Bull. Ibaraki Univ. (5): 13 (taxonomy), 19 (key), figs. 29-34..

 

facialis du Buysson

Belonogaster facialis du Buysson, 1908, Bull. Soc. Entomol. France 1908: 65, female -- gAfrique orientale anglaise: Boura, Wa-Taïtah [Kenya] (lectotype Paris); also from Senegal; and Congo; 1909, Ann. Soc. Entomol. France 78 (2): 215, 238; 1914, in Voy. Alluaud et Jeannel Afr. Or., Rés. Sci., Ins. Hym. 3, Vesp.: 161. -- von Schulthess, 1913, Ark. Zool. 8 (17): 14 (syn.: sex-maculatus Cameron). -- Bequaert, 1918, Bull. Am. Mus. Nat. Hist. 39: 330 (cat.). -- Richards, 1982, Bull. Br. Mus. (Nat. Hist.) Entomol. 44: 34, 44 (key), 80, figs. 51--53 (designation of lectotype; distr.).

Distribution: Senegal; Congo; Kenya; Uganda; Tanzania (including Zanzibar); Zambia; Malawi; Mozambique; South Africa (Kwazulu-Natal).

 

fanemitra Hensen and Blommers

Belonogaster fanemitra Hensen and Blommers, 1987, Tijdschr. Entomol. 130: 14 (key), 22, figs. 20, 38, 39, male, female -- gMadagascar, Sambirano, Manongarivo, 1150 mh (holotype female Paris); also from other locality in Madagascar.

Distribution: Madagascar.

 

ferruginea Richards

Belonogaster ferruginea Richards, 1982, Bull. Br. Mus. (Nat. Hist.) Entomol. 44: 36 (key), 52, female -- gGrande Comore: La Grille (Guiri), 850--900 mh (Paris).

Distribution: Comoros.

 

filiformis (de Saussure)

Raphigaster filiformis de Saussure, 1853, Ét. Fam. Vesp. 2: 18, pl. II fig. 4, female -- gDjidda en Arabieh (lectotype Paris). -- Smith, 1857, Cat. Hym. Br. Mus. 5: 94 (syn. of Belonogaster macilentus (Fabricius)). -- Dalla Torre, 1894, Cat. Hym. 9: 114 (Rhaphigaster[!]). -- Richards, 1982, Bull. Br. Mus. (Nat. Hist.) Entomol. 44: 100 (designation of lectotype).

Belonogaster filiformis; Kohl, 1894 (1893), Ann. Naturhist. Hofmus. Wien 9: 335. -- Dalla Torre, 1894, Cat. Hym. 9: 114. -- Gribodo, 1895, Mem. R. Accad. Sci. Bologna (5) 5: 335. -- Dalla Torre, 1904, Genera Insectorum 19: 80 (cat.). -- du Buysson, 1909, Ann. Soc. Entomol. France 78 (2): 216--217, 257. -- Bequaert, 1918, Bull. Am. Mus. Nat. Hist. 39: 330 (cat.; gperhaps a variety of B. griseus (Fabricius)h). -- Richards, 1982, Bull. Br. Mus. (Nat. Hist.) Entomol. 44: 40, 45 (key), 100, fig. 78 (distr.); 1985 (1984), Fauna Saudi Arabia 6: 436, 437 (key), 438 (distr.).

Distribution: Saudi Arabia.

 

filiventris (de Saussure)

Raphigaster filiventris de Saussure, 1853, Ét. Fam. Vesp. 2: 16, pl. 2 fig. 5, female -- gLe Sénégal?h (Paris). -- Dalla Torre, 1894, Cat. Hym. 9: 114 (Rhaphigaster[!]).

Belonogaster filiventris; Smith, 1857, Cat. Hym. Br. Mus. 5: 94 (cat.). -- Dalla Torre, 1894, Cat. Hym. 9: 114 (cat.), 116 (?syn. of B. longistyla). -- Kohl, 1894 (1893), Ann. Naturhist. Hofmus. Wien 9: 335. -- Gribodo, 1894, in Emery et al., Mem. R. Accad. Sci. Bologna (5) 4: 123; 1895, Mem. R. Accad. Sci. Bologna (5) 5: 335. -- Bingham, 1903, Ann. Mag. Nat. Hist. (7) 12: 47, 68. -- Dalla Torre, 1904, Genera Insectorum 19: 80 (cat.). -- du Buysson, 1909, Ann. Soc. Entomol. France 78 (2): 216, 218, 241, 264, pl. 4 fig. 1, pl. 6 fig. 3 (male; ethology); 1914, in Voy. Alluaud et Jeannel Afr. Or., Rés. Sci., Ins. Hym. 3, Vesp.: 162. -- Bequaert, 1918, Bull. Am. Mus. Nat. Hist. 39: 331 (cat.; syn.: B. braunsii Kohl, B. kohli Schulz, B. gracilis Cameron). -- Salt, 1927, Psyche 34: 185 (distr.; record of stylopized specimen). -- von Schulthess, 1928, Senckenbergiana 10 (3/4): 96 (distr.). -- Salt and Bequaert, 1929, Psyche 36: 262 (record of stylopized specimen). -- Bequaert, 1938, Explor. Parcs Nation. Congo Belge Miss. G. F. de Witte (1933--1935) 11: 9 (distr.). -- Giordani Soika, 1961, S. Afr. Anim. Life 8: 450 (distr.). -- Marino Piccioli, 1968, Monit. Zool. Ital. (n. ser.) 2, Suppl.: 203--206 (ethology). -- Richards, 1982, Bull. Br. Mus. (Nat. Hist.) Entomol. 44: 36, 42 (key), 75, figs. 45--47 (syn.: B. gracilis Cameron, B. sexmaculatus Cameron, B. buyssoni Meade-Waldo; distr.).

Belonogaster gracilis Cameron, 1910, in Sjöstedt, Wiss. Ergebn. Schwed. Zool. Exp. Kilimandjaro 2 (8), Hym. 6: 173, female -- gKilimandjaro: Kibonoto, cultivated zone, 1300--1900 mh [Tanzania] (lectotype Stockholm). -- von Schulthess, 1913, Ark. Zool. 8 (17): 14 (syn. of B. filiventris (de Saussure)). -- Richards, 1982, Bull. Br. Mus. (Nat. Hist.) Entomol. 44: 76 (designation of lectotype; marked as gSyn. n.h).

Belonogaster sexmaculatus Cameron, 1910, in Sjöstedt, Wiss. Ergebn. Schwed. Zool. Exp. Kilimandjaro 2 (8), Hym. 6: 174, male -- gKilimandjaro: Kibonoto, cultivated zoneh [Tanzania] (Stockholm). -- von Schulthess, 1913, Ark. Zool. 8 (17): 14 (syn. of B. facialis du Buysson). -- Richards, 1982, Bull. Br. Mus. (Nat. Hist.) Entomol. 44: 76 (syn. of B. filiventris (de Saussure)).

Belonogaster buyssoni Meade-Waldo , 1911, Ann. Mag. Nat. Hist. (8) 7: 99, female -- gIganga Busoga, S. Nigeriah (London). -- Bequaert, 1918, Bull. Am. Mus. Nat. Hist. 39: 329 (cat.; gProbably a synonym of B. pusillus Kohlh); 1928, Ann. Mag. Nat. Hist. (10) 2: 173 (the type is from Uganda, not S. Nigeria, as stated by error in the original description). -- Richards, 1982, Bull. Br. Mus. (Nat. Hist.) Entomol. 44: 76 (syn. of B. filiventris (de Saussure)).

Distribution: Liberia; Nigeria; Cameroon; Zaire; Uganda; Kenya; Tanzania; Angola; Zambia; Malawi; Zimbabwe; Mozambique; South Africa (Kwazulu-Natal, Transvaal, Western Cape).

 

flava Richards

Belonogaster flava Richards, 1982, Bull. Br. Mus. (Nat. Hist.) Entomol. 44: 36 (key), 75, figs. 43, 44, female -- gUganda: Budongo Forest, Unyoro, 3400 ft [1040 m]h (London); also from Cameroon.

Distribution: Uganda; Cameroon.

 

freyi du Buysson

Belonogaster freyi du Buysson, 1909, Ann. Soc. Entomol. France 78 (2): 217 (key), 255, female -- gAfrique australeh (holotype Genève). -- Bequaert, 1918, Bull. Am. Mus. Nat. Hist. 39: 331 (cat.). -- Richards, 1982, Bull. Br. Mus. (Nat. Hist.) Entomol. 44: 39, 44 (key), 69, fig. 37 (syn.: indicus var. claripennis; distr.). — Kojima, 2001a, Nat. Hist. Bull. Ibaraki Univ. (5): 20.

Belonogaster indicus var. claripennis du Buysson, 1909, Ann. Soc. Entomol. France 78 (2): 217, 219, 259, pl. 4 fig. 9, female, male -- gAfrique orientale allemande: Kigonsera; Ougandah [Tanzania] (holotype female Paris). -- Bequaert, 1918, Bull. Am. Mus. Nat. Hist. 39: 333 (cat.). -- Richards, 1982, Bull. Br. Mus. (Nat. Hist.) Entomol. 44: 69 (syn. of B. freyi du Buysson).

Distribution: Zaire; Kenya; Tanzania; Zambia; Malawi; Zimbabwe; Botswana; South Africa (Kwazulu-Natal, Free State, Transvaal).

 

fuscipennis du Buysson

Belonogaster griseus var. fuscipennis du Buysson, 1909, Ann. Soc. Entomol. France 78 (2): 251, 264, female, male -- gCongo français: Haute Sangah (lectotype female Paris). -- Bequaert, 1918, Bull. Am. Mus. Nat. Hist. 39: 332 (cat.). -- Schouteden, 1919, Rev. Zool. Afr. 6: 186 (distr.). -- von Schulthess, 1928, Senckenbergiana 10 (3/4): 96 (distr.). -- Richards, 1982, Bull. Br. Mus. (Nat. Hist.) Entomol. 44: 86 (designation of lectotype).

Belonogaster fuscipennis; Richards, 1982, Bull. Br. Mus. (Nat. Hist.) Entomol. 44: 39 (key), 86 (distr.).

Distribution: Congo; Zaire; Sudan.

 

grisea (Fabricius)

Vespa grisea Fabricius, 1775, Syst. Entomol.: 372 -- gSierra Leon Africaeh (holotype female London); 1781, Spec. Insectorum 1: 468; 1787, Mant. Insectorum 1: 293. -- Gmelin, 1790, Syst. Nat., ed 13, 1 (5): 2753. -- Olivier, 1792, Encycl. Méthod. Insectes 6: 672. -- Fabricius, 1793, Entomol. Syst. 2: 279.

Sphex grisea; Christ, 1791, Naturg. Insekten.: 313.

Eumenes grisea; Fabricius, 1804, Syst. Piez.: 286.

Raphigaster rufipennis; de Saussure, 1853, Ét. Fam. Vesp. 2: 15, pl. 2 fig. 6, (female) [Misidentification of Sphex rufipennis DeGeer]. -- Dalla Torre, 1894, Cat. Hym. 9: 115 (Rhaphigaster [!]; syn of B. grisea).

Belonogaster rufipennis; Smith, 1857, Cat. Hym. Br. Mus. 5: 93. -- Saunders, 1872, Trans. R. Entomol. Soc. London 1872: 36 (record of stylopized specimen). -- Radoszkowski, 1876, Horae Soc. Entomol. Ross. 12, 2: 141; 1881, J. Sci. Math. Phys. Nat. Acad. Lisbon 8 (31): 203. -- Fox, 1891, Entomol. News 2: 42. -- Schletterer, 1891, Ann. Soc. Entomol. Belg. 35: 28. -- Distant, 1892, A Naturalist in the Transvaal: 210. -- Gribodo, 1894, in Emery et al., Mem. R. Accad. Sci. Bologna (5) 4: 123 (rufipennis?). -- Dalla Torre, 1894, Cat. Hym. 9: 115 (syn. of B. grisea). -- Gribodo, 1895, Mem. R. Accad. Sci. Bologna (5) 5: 335. -- du Buysson, 1898, Ann. Soc. Entomol. France 66, 2--3: 362. [Misidentification of Sphex rufipennis DeGeer]. -- Dalla Torre, 1904, Genera Insectorum 19: 80 (syn. of B. grisea). -- Salt and Bequaert, 1929, Psyche 36: 263 (syn. of B. griseus (Fabr.); record of stylopized specimen).

Belonogaster griseus; Smith, 1857, Cat. Hym. Br. Mus. 5: 94 (cat.). -- Saunders, 1872, Trans. R. Entomol. Soc. London 1872: 36 (record of stylopized specimen). -- Radoszkowski, 1881, J. Sci. Math. Phys. Nat. Acad. Lisboa 8 (31): 203. -- Kohl, 1894 (1893), Ann. Naturhist. Hofmus. Wien 9: 335. -- Gribodo, 1894, in Emery et al., Mem. R. Accad. Sci. Bologna (5) 4: 123. -- Dalla Torre, 1894, Cat. Hym. 9: 115 (grisea; cat.; syn.: Vespa macilenta Fabricius, B. rufipennis de Saussure). -- Magretti, 1898, Ann. Mus. Civ. Genova (2), 19: 37. -- Bingham, 1903, Ann. Mag. Nat. Hist. (7) 12: 47. -- Dalla Torre, 1904, Genera Insectorum 19: 80 (grisea; cat.; as 1894). -- von Schulthess, 1907, Voeltzkow, Reise in Ostafrika 1903--1905, Wiss. Ergebn. 2 (2): 64. -- du Buysson, 1909, Ann. Soc. Entomol. France 78 (2): 217, 218, 247, 264, pl. 2 figs. 11--12, pl. 4 fig. 8, pl. 5 fig. 3 (male; ethology). -- Cameron, 1910, Ann. Transvaal Mus. 2, 3: 155. -- Roubaud, 1910, C. R. Acad. Sci. Paris 151: 553 (ethology). -- von Schulthess, 1912, Wiss. Ergebn. D. Z. Afrik. Exp., (1907--08), 4, Lf. 10: 291; 1912, Soc. Entomol. 27: 41 (compared to B. atratus n. sp.); 1913, Mitt. Zool. Mus. Berlin 6 (3): 339; 1914, Dtsch. Entomol. Z. 1914: 288. -- du Buysson, 1914, in Voy. Alluaud et Jeannel Afr. Or., Rés. Sci., Ins. Hym. 3, Vesp.: 162. -- Roubaud, 1916, Ann. Sci. Nat. Zool. (10), 1, 1: 110, 139 (ethology). -- Bequaert, 1918, Bull. Am. Mus. Nat. Hist. 39: 236 (key), 237, 331 (cat.; distr.; syn.: B. macilenta (Fabricius), B. linearis (Olivier), B. rufipennis (de Saussure), B. pictus Kohl, B. erythrospilus Cameron, B. fulvipennis de Saussure). -- Schouteden, 1919, Rev. Zool. Afr. 6: 185 (distr.). -- von Schulthess, 1922, Dtsch. Entomol. Z.: 400 (distr.); 1928, Senckenbergiana 10 (3/4): 96 (distr.); 1929, Rev. Zool. Bot. Afr. 17 (2): 187 (distr.). -- Salt and Bequaert, 1929, Psyche 36: 262 (record of stylopized specimen). -- Bequaert, 1933, Ann. Mag. Nat. Hist. 12: 117 (distr.). -- Giordani Soika, 1935, Mem. Estud. Mus. Zool. Univ. Coimbra (1) 82: 13 (distr.). -- Bequaert, 1938, Explor. Parcs Nation. Congo Belge Miss. G. F. de Witte (1933--1935) 11: 8, 9, 10 (distr.). -- Guiglia, 1938, Ann. Mus. Civ. Stor. Nat. Genova 58: 182 (distr.); 1939, Mem. Soc. Entomol. Ital. 17: 192 (distr.); 1940, Boll. Lab. Zool. Gen. Agrar. Fac. Agrar. Portici 31: 279 (var.?). -- FitzGerald, 1940, Proc. R. Entomol. Soc. London Ser. A Gen. Entomol. 15: 35 (distr.). -- Berland and Grassé, 1951, in Grassé, Traité Zool. 10 (2): fig. 1008--1. -- Benoit, 1956, Ann. Mus. R. Congo Belge 8 (Sci. Zool.) 51: 552 (distr.). -- Marino Piccioli, 1968, Monit. Zool. Ital. (n. ser.) 2, Suppl.: 203 (ethology). -- Marino Piccioli and Pardi, 1970, Monit. Zool. Ital. (n. ser.) 3, Suppl.: 197 (ethology.). -- Pardi and Marino Piccioli, 1970, Monit. Zool. Ital. (n. ser.) 3, Suppl. (11): 235 (ethology). -- Pardi, 1977, Boll. Inst. Entomol. Univ. Bologna 33: 281 (ethology). -- Marino Piccioli and Pardi, 1978, Monit. Zool. Ital. (n. ser.) 10, Suppl.: 179 (ethology). -- Pardi and Marino Piccioli, 1981, Monit. Zool. Ital. (n. ser.) 14, Suppl. (9): 131 (ethology). -- Richards, 1982, Bull. Br. Mus. (Nat. Hist.) Entomol. 44: 39, 45 (key), 87, fig. 62 (grisea; syn.: B. pictus Kohl, B. griseus var. pallens du Buysson, ?B. braunsii Kohl, B. erythrospilus Cameron; distr.). -- Jeanne et al., 1983, Zoomorphologie 103: 155, figs. 7, 12, 18 (grisea; morphology). -- Spradbery, 1991, in Ross and Matthews, Soc. Biol. Wasps: 354, 355, 357 (grisea; colony population). -- Jeanne, 1991, in Ross and Matthews, Soc. Biol. Wasps: 393 (gresea; ethology ). -- Ross and Carpenter, 1991, in Ross and Matthews, Soc. Biol. Wasps: 463 (grisea biology). -- Downing, 1991, in Ross and Matthews, Soc. Biol. Wasps: 549 (grisea; morphology). -- Itô, 1993, Behav. Soc. Evol. Wasps: 20, 40, 104, 131, 132 (ethology).

Belonogaster fulvipennis [!]; de Saussure, 1890, in Grandidier, Hist. Madagascar 20, Hym. 1: 87 footnote, 90 [misspelling of rufipennis]. -- Kohl, 1894 (1893), Ann. Naturhist. Hofmus. Wien 9: 335. -- Dalla Torre, 1894, Cat. Hym. 9: 115 (cat.); 1904, Genera Insectorum 19: 80 (cat.).

Belonogaster pictus Kohl, 1894 (1893), Ann. Naturhist. Hofmus. Wien 9: 320, 323 (key), 324, 337, pl. 16 fig. 118, pl. 18 figs. 142, 146, 152, 156, female, male -- gCamerunh; (lectotype female Wien); also from other localities. -- Dalla Torre, 1894, Cat. Hym. 9: 116 (picta; cat); 1904, Genera Insectorum 19: 80 (picta; cat.). -- Tullgren, 1904, Ark. Zool. 1: 454 (Belenogaster [!]). -- Cameron, 1906, Trans. S. Afr. Philos. Soc. 16 (4): 327. -- Schulz, 1906, Spolia Hym.: 321. -- Maidl, 1913, Sitz. Ber. K. Akad. Wiss. Wien, Math. Naturw. Kl. 122 (4), Abt. 1: 560. -- Zavattari, 1909, Il Ruwenzori, Parte Scientif. 1, Zool.: 211. -- Richards, 1982, Bull. Br. Mus. (Nat. Hist.) Entomol. 44: 88 (designation of lectotype).

?Belonogaster braunsii Kohl, 1894 (1893), Ann. Naturhist. Hofmus. Wien 9: 323 (key), 331, 334, female -- gSüdafrika: Port Natalh (type depository unknown). -- Dalla Torre, 1894, Cat. Hym. 9: 114 (cat.); 1904, Genera Insectorum 19: 80 (braunsi [!]; cat.). -- Bequaert, 1918, Bull. Am. Mus. Nat. Hist. 39: 331 (syn. of filiventris (de Saussure)). -- Giordani Soika, 1961, S. Afr. Anim. Life 8: 450 (braunsi [!]; syn. of filiventris (de Saussure)). -- Richards, 1982, Bull. Br. Mus. (Nat. Hist.) Entomol. 44: 88 (?syn. of grisea).

Belonogaster griseus var. pallens du Buysson, 1909, Ann. Soc. Entomol. France 78 (2): 216, 250, 265, female, male -- gDakarh (lectotype female Paris). -- Bequaert, 1918, Bull. Am. Mus. Nat. Hist. 39: 332 (cat.). -- Schouteden, 1919, Rev. Zool. Afr. 6: 186 (distr.). -- Richards, 1982, Bull. Br. Mus. (Nat. Hist.) Entomol. 44: 88 (designation of lectotype).

Belonogaster erythrospilus Cameron, 1910, in Sjöstedt, Wiss. Ergebn. Schwed. Zool. Exp. Kilimandjaro 2 (8), Hym. 6: 172, female, male -- gMeru low lands, Ngare na nyukih [Tanzania] (lectotype male Stockholm); also from Kilimandjaro and Usambara. -- von Schulthess, 1913, Ark. Zool. 8 (17): 14 (syn. of grisea). -- Bequaert, 1933, Ann. Mag. Nat. Hist. 12: 117 (perhaps color form of griseus). -- Richards, 1982, Bull. Br. Mus. (Nat. Hist.) Entomol. 44: 88 (designation of lectotype, listed as both female and male; ?syn. of B. grisea).

Distribution: Senegal; Guinea-Bissau; Guinea; Sierra Leone; Liberia; Ghana; Nigeria; Chad; Sudan; Cameroon; Bioko; Gabon; Central African Republic; Congo; Zaire; Uganda; Rwanda; Burundi; Kenya; Tanzania (including Zanzibar); Angola; Zambia; Malawi; Zimbabwe; Mozambique; Botswana; South Africa (Transvaal, Kwazulu-Natal, Western Cape); Swaziland.

 

guerini de Saussure

Raphigaster guerini de Saussure, 1853, Ét. Fam. Vesp. 2: 17, pl. 2 fig. 3, female -- gMadagascarh (Genova). -- Dalla Torre, 1894, Cat. Hym. 9: 115 (Rhaphigaster [!]).

Belonogaster guerini; Smith, 1857, Cat. Hym. Br. Mus. 5: 91 (cat.). -- de Saussure, 1890, in Grandidier, Hist. Madagascar 20, Hym. 1: 89 (key), 91 (male). -- Kohl, 1894 (1893), Ann. Naturhist. Hofmus. Wien 9: 335. -- Dalla Torre, 1894, Cat. Hym. 9: 115 (guerinii [!]; cat.). -- Gribodo, 1894, in Emery et al., Mem. R. Accad. Sci. Bologna (5) 4: 123. -- von Schulthess, 1899, Bull. Soc. Vaudoise Sci. Nat. (4), 35: 269 (distr.). -- de Saussure, 1900, Abh. Senckenb. Naturforsch. Ges. 26 (2): 207 (key), 208. -- Dalla Torre, 1904, Genera Insectorum 19: 80 (cat.). -- von Schulthess, 1907, Voeltzkow, Reise in Ostafrika 1903--1905, Wiss. Ergebn. 2 (2): 65. -- du Buysson, 1909, Ann. Soc. Entomol. France 78 (2): 214, 225, pl. 5 figs. 8--14 (ethology; distr. erroneous). -- Bequaert, 1918, Bull. Am. Mus. Nat. Hist. 39: 332 (cat.; distr. erroneous). -- Salt and Bequaert, 1929, Psyche 36: 262 (record of stylopized specimen). -- Richards, 1982, Bull. Br. Mus. (Nat. Hist.) Entomol. 44: 46 (key), 102, fig. 81 (distr.). -- Hensen and Blommers, 1987, Tijdschr. Entomol. 130: 12 (key), 23, fig. 44 (distr.; nest).

Distribution: Madagascar.

 

guichardi Richards

Belonogaster guichardi Richards, 1982, Bull. Br. Mus. (Nat. Hist.) Entomol. 44: 40, 44 (key), 94, fig. 70, male, female -- gOman: Dhofar, Ayun Pools, 700 mh (holotype male London); also from two other localities in Oman; 1985 (1984), Fauna Saudi Arabia 6: 436, 437 (key), 438 (Oman).

Distribution: Oman.

 

hildebrandti de Saussure

Belonogaster hildebrandti de Saussure, 1890, in Grandidier, Hist. Madagascar 20, Hym. 1: 89 (key), 95, pl. 17 fig. 11, female, male -- gMadagascar, région centraleh (lectotype female Berlin). -- Kohl, 1894 (1893), Ann. Naturhist. Hofmus. Wien 9: 335. -- Dalla Torre, 1894, Cat. Hym. 9: 115 (hildebrandtii [!]; cat.). -- de Saussure, 1900, Abh. Senckenb. Naturforsch. Ges. 26 (2): 207 (key). -- Dalla Torre, 1904, Genera Insectorum 19: 80 (cat.). -- du Buysson, 1909, Ann. Soc. Entomol. France 78 (2): 214, 232, pl. 4, fig. 5, pl. 5 figs. 5, 7 (ethology). -- Bequaert, 1918, Bull. Am. Mus. Nat. Hist. 39: 333 (cat.). -- Richards, 1982, Bull. Br. Mus. (Nat. Hist.) Entomol. 44: 47 (key), 108, fig. 90 (designation of lectotype; distr.). -- Hensen and Blommers, 1987, Tijdschr. Entomol. 130: 14 (key), 23 (sexes associated by Richards doubtfully conspecific).

Distribution: Madagascar.

 

hirsuta Richards

Belonogaster hirsuta Richards, 1982, Bull. Br. Mus. (Nat. Hist.) Entomol. 44: 41 (key), 55, male -- gTanzania: Mahali peninsula, Kungwe Camph (London).

Distribution: Tanzania.

 

indica de Saussure

Raphigaster indicus de Saussure, 1853, Ét. Fam. Vesp. 2: 17, female -- gLes Indes Orientales. Bombayh (Paris). --  Dalla Torre, 1894, Cat. Hy. 9: 115 (Rhaphigaster [!]).

Belonogaster indicus; Smith, 1857, Cat. Hym. Br. Mus. 5: 94 (cat.); 1871, J. Proc. Linn. Soc. Zool. 11: 378 (cat.). -- de Saussure, 1890, in Grandidier, Hist. Madagascar 20, Hym. 1: 87. -- Kohl, 1894 (1893), Ann. Naturhist. Hofmus. Wien 9: 335. -- Gribodo, 1894, in Emery et al., Mem. R. Accad. Sci. Bologna (5) 4: 123. -- Dalla Torre, 1894, Cat. Hym. 9: 115 (indica; cat.). -- Gribodo, 1895, Mem. R. Accad. Sci. Bologna (5) 5: 335 -- Bingham, 1897, Fauna Br. India, Hym. 1: 382, fig. 114 (female, male; distr.). -- Dalla Torre, 1904, Genera Insectorum 19: 80 (indica; cat.). -- du Buysson, 1909, Ann. Soc. Entomol. France 78 (2): 214, 258 (female). -- Bequaert, 1918, Bull. Am. Mus. Nat. Hist. 39: 333 (cat.). -- Berland and Grassé, 1951, in Grassé, Traité Zool. 10 (2): 1152. -- Richards, 1982, Bull. Br. Mus. (Nat. Hist.) Entomol. 44: 31, 35 (key), 58 (indica; redescription of the type). -- Das and Gupta, 1984 (1983), Orient. Insects 17: 432 (indica; cat.); 1989, Orient. Insects Monogr. 11: 186 (key), 187, map 30 (indica; distr.).

Distribution: India.

 

jordani Richards

Belonogaster jordani Richards, 1982, Bull. Br. Mus. (Nat. Hist.) Entomol. 44: 38 (key), 83, female -- gAngola: Quirimboh (London).

Distribution: Angola.

 

juncea colonialis Kohl

Belonogaster colonialis Kohl, 1894 (1893), Ann. Naturhist. Hofmus. Wien 9: 320, 323 (key), 335, pl. 15 figs. 72, 84, pl. 17 figs. 132, 139, female, male -- gOstafrika: Dar-es-Salaamh (lectotype male Wien). -- Dalla Torre, 1894, Cat. Hym. 9: 114; 1904, Genera Insectorum 19: 80 (cat.). -- Fox, 1896, Proc. Acad. Nat. Sci. Philadelphia 48: 555. -- Stadelmann, 1898, Dtsch. Ost-Afrika 4, Hym.: 35. -- Bingham, 1909, Trans. Zool. Soc. London 19 (2): 181. -- Zavattari, 1909, Il Ruwenzori, Parte Scientif. 1, Zool.: 210. -- Bequaert, 1918, Bull. Am. Mus. Nat. Hist. 39: 334 (syn. of B. juncea). -- Giordani Soika, 1961, S. Afr. Anim. Life 8: 451. -- Richards, 1982, Bull. Br. Mus. (Nat. Hist.) Entomol. 44: 48 (designation of lectotype).

Belonogaster junceus var. colonialis; Bequaert, 1938, Explor. Parcs Nation. Congo Belge Miss. G. F. de Witte (1933--1935) 11: 9 (distr.).

Belonogaster juncea colonialis; Richards, 1982, Bull. Br. Mus. (Nat. Hist.) Entomol. 44: 35, 44 (key), 48 (distr.). -- Keeping and Crewe, 1983, J. Entomol. Soc. South. Africa 46: 309 (ethology.). -- Kojima and Keeping, 1984, J. Entomol. Soc. South Africa 48: 234--237 (larva).

Distribution: Congo; Zaire; Uganda; Kenya; Tanzania (including Zanzibar); Angola; Zambia; Malawi; Zimbabwe; Mozambique; South Africa (Transvaal, Kwazulu-Natal).

 

juncea juncea (Fabricius)

Vespa juncea Fabricius, 1781, Spec. Insectorum 1: 468 -- gAfrica aequinoctialih (holotype female London); 1787, Mant. Insectorum 1: 293. -- Gmelin, 1790, Syst. Nat., ed. 13, 1 (5): 2754. -- Christ, 1791, Naturg. Inseckten 6: 673. -- Olivier, 1792, Encycl. Méthod. Insectes 6: 673.

?Vespa guineensis Fabricius, 1793, Entomol. Syst. 2: 277 -- gGuineah (holotype female Kiel). -- Dalla Torre, 1894, Cat. Hym. 9: 115 (syn of B. juncea).

Vespa cinerea Fabricius, 1793, Entomol. Syst. 2: 279 -- gAfrica aequinoctialih (?London, Kiel). -- Jurine, 1807, Nouv. Méth. Class. Hym.: 172 (cinereus; female, male). -- Dalla Torre, 1894, Cat. Hym. 9: 165 (syn. of B. juncea).

Zethus guineensis; Fabricius, 1804, Syst. Piez.: 283.

Zethus cinereus; Fabricius, 1804, Syst. Piez.: 283.

Polistes cinerea; Latreille, 1809, Gen. Crusta. Insestes 4: 142.

Raphigaster junceus; de Saussure, 1853, Ét. Fam. Vesp. 2: 14, pl. 2 fig. 2 (female, male). -- Dalla Torre, 1894, Cat. Hym. 9: 115 (Rhaphigaster [!]).

Raphigaster guineensis; Smith, 1856, Trans. R. Entomol. Soc. London (2) 3, Proc.: 129 (ethology). -- Dalla Torre, 1894, Cat. Hym. 9: 115 (Rhaphigaster [!]; syn of B. juncea (Fabr.)).

Belonogaster junceus; Smith, 1857, Cat. Hym. Br. Mus. 5: 93 (cat.; syn.: Vespa juncea Fabricius, Vespa Guineensis Fabricius, Vespa cinerea Fabricius); 1859, Trans. R. Entomol. Soc. London (2) 5 (3): 130 (record of stylopized specimen). -- Gerstaecker, 1862, in Peters, Naturwiss. Reise Mossambique, Hym.: 468; 1871, Arch. Naturg. 38, 1: 351. -- Walker, 1871, List Hym. Egypt: 39. -- Gerstaecker, 1873, in v. d. Denkenfs Reisen in Ost-Afrika 3, 2, Gliederth.: 324. -- Ritsema, 1874, Tijdschr. Entomol. 17: 177, 201, 210. -- Magretti, 1884, Ann. Mus. Civ. Genova 21: 599 (ethology). -- Gribodo, 1884, Ann. Mus. Civ. Genova 21: 289. -- Schletterer, 1891, Ann. Soc. Entomol. Belgique 35: 28. -- von Schulthess, 1893, Entomol. Nachr. 19: 18. -- Kohl, 1893 (1892), Jahrb. Hamburg. Wiss. Anst. 10 (2): 186; 1894, Ann. Naturhist. Hofmus. Wien 9: 336. -- Gribodo, 1894, in Emery et al., Mem. R. Accad. Sci. Bologna (5) 4: 123. -- Dalla Torre, 1894, Cat. Hym. 9: 115 (juncea; cat.; syn.: Vespa juncea Fabricius, V. Guineensis Fabricius, V. cinerea Fabricius). -- Gribodo, 1895, Mem. R. Accad. Sci. Bologna (5) 5: 334. -- Magretti, 1898, Ann. Mus. Civ. Genova (2), 19: 36. -- Stadelmann, 1898, Dtsch. Ost-Afrika 4, Hym.: 35. -- Bingham, 1903, Ann Mag. Nat. Hist. (7) 12: 48. -- Dalla Torre, 1904, Genera Insectorum 19: 80, pl. 5, fig. 6 (juncea; cat.; as in 1894). -- Tullgren, 1904, Ark. Zool. 1: 453 (Belenogaster [!]; ethology). -- du Buysson, 1908, in A. Chevalier, LfAfrique Centrale Française: 707; 1909, Ann. Soc. Entomol. France 78 (2): 213, 220, 263, pl. 2 figs. 1--7, pl. 3 figs. 1--7, pl. 4 figs. 3, 12--13, pl. 5 fig. 4, pl. 7 fig. 2 (female, male; ethology). -- Bingham, 1909, Trans. Zool. Soc. London 19, 2: 181. -- Roubaud, 1910, C. R. Acad. Sci. Paris 151: 553--556 (ethology). -- Cameron, 1910, Ann. Transvaal Mus. 2, 3: 156; 1910, in Sjöstedt, Wiss. Ergebn. Schwed. Zool. Exp. Kilimandjaro 2 (8), Hym. 6: 171. -- Schulz, 1911, Zool. Ann. Würzburg 4: 200. -- von Schulthess, 1912, Wiss. Ergebn. D. Z. Afrik. Exp., (1907--08), 4, Lf. 10: 291; 1912, Soc. Entomol. 27: 41; 1913, Mitt. Zool. Mus. Berlin 6 (3): 337; 1913, Ark. Zool. 8 (17): 14. -- Maidl, 1913, Sitz. Ber. K. Akad. Wiss. Wien, Math. Naturw. Kl. 122, Abt. 1 (4): 560. -- von Schulthess, 1914, Dtsch. Entomol. Z. 1914: 288. -- du Buysson, 1914, in Voy. Alluaud et Jeannel Afr. Or., Rés. Sci., Ins. Hym. 3, Vesp.: 160. -- Roubaud, 1916, Ann. Sci. Nat. Zool. (10) 1 (1): 110--139 (ethology). -- Bequaert, 1918, Bull. Am. Mus. Nat. Hist. 39: 236 (key), 333 (syns: Vespa cinerea Fabricius, Raphigaster guineensis (Smith), B. colonialis Kohl; cat; distr.). -- Schouteden, 1919, Rev. Zool. Afr. 6: 184 (distr.; misidentified by von Schulthess as abyssinicus). -- von Schulthess, 1922, Denkschr. Akad. Wiss. Wien, Math. Naturh. Kl. 98 (6): 96 (distr.). -- Salt, 1927, Psyche 34: 185 (distr.; record of stylopized specimen). -- von Schulthess, 1928, Senckenbergiana 10 (3/4): 95 (distr.); 1929, Rev. Zool. Bot. Afr. 17 (2): 186 (distr.). -- Salt and Bequaert, 1929, Psyche 36: 262 (record of stylopized specimen). -- Guiglia, 1932 (1931), Mem. Soc. Entomol. Ital. 10: 118 (distr.). -- Bequaert, 1938, Explor. Parcs Nation. Congo Belge Miss. G. F. de Witte (1933--1935) 11: 8, 9, 10 (distr.). -- Guiglia, 1939, Mem. Soc. Entomol. Ital. 18: 88; 1940, Boll. Lab. Zool. Gen. Agrar. Fac. Agrar. Portici 31: 279 (distr.). -- Giordani Soika, 1940 (1939), Mem. Soc. Entomol. Ital. 17: 99 (distr.); 1952 (1951), Riv. Biol. Colon. 11: 88 (distr.); 1961, S. Afr. Anim. Life 8: 451 (Zambia; Zimbabwe; South Africa: Transvaal). -- Richards, 1969, Mem. Soc. Entomol. Ital. 48: 87--93 (ethology). -- Spradbery, 1991, in Ross and Matthews, Soc. Biol. Wasps: 354, 355 (juncea; colony population). -- Ross and Carpenter, 1991, in Ross and Matthews, Soc. Biol. Wasps: 463 (juncea; biology). -- Itô, 1993, Behav. Soc. Evol. Wasps: 20 (juncea), 132 (juncus [!]; ethology).

Belonogaster juncea juncea; Richards, 1982, Bull. Br. Mus. (Nat. Hist.) Entomol. 44: 35, 44 (key), 47, figs. 1--5 (?syn.: Vespa guineensis Fabricius; distr.). -- Das and Gupta, 1984 (1983), Orient. Insects 17: 433 (cat.). -- Richards, 1985 (1984), Fauna Saudi Arabia 6: 426 (j. junceus), 436, 437, 438, fig. 10 (key; distr.; list). -- Das and Gupta, 1989, Orient. Insects Monogr. 11: 186 (key), 188, fig. 32, map 30. -- Dejean and Turillazzi, 1992, Trop. Zool. 5: 237--247 (trophobiosis with homopterans). -- Francescato et al., 1994, Ethol. Ecol. Evol. Spec. Issue 3: 53--56 (ethology). -- Tindo et al., 1994, in Lenoir et al., Insect. Soc.: 532 (ethology). -- Tindo et al., 1997, Insestes Sociaux 44: 365 (ethology). -- Tindo et al., 1997, J. Insect Behav. 10: 571 (ethilogy). -- Tindo et al., 1997, Sociobiology 29: 255 (ethology).

Distribution: Libya; Senegal; Gambia; Mali; Chad; Sudan; Guinea; Sierra Leone; Liberia; Côte dfIvoire; Ghana, Togo; Benin Republic; Nigeria; Cameroon; Eritrea; Ethiopia; Gabon; Congo; Zaire; Uganda; Rwanda; Kenya; Tanzania; Angola; Zambia; Malawi; Mozambique; South Africa; Saudi Arabia; India.

 

kelnerpillautae Richards

Belonogaster kelnerpillautae Richards, 1982, Bull. Br. Mus. (Nat. Hist.) Entomol. 44: 36 (key), 77, female -- gCongo: Dimonikah (Paris); also from other locality in Congo.

Distribution: Congo.

 

kohli Schulz

Belonogaster kohli Schulz, 1906, Spolia Hym.: 322, female -- gInsel Fernando Poh [Malabo (Santa Isabel)] (London). -- Bequaert, 1918, Bull. Am. Mus. Nat. Hist. 39: 331 (syn. of B. filiventris (de Saussure)). -- Richards, 1982, Bull. Br. Mus. (Nat. Hist.) Entomol. 44: 35 (key), 53, fig. 9 (Bioko, Congo, Uganda).

Distribution: Bioko; Congo; Uganda.

 

lateritia Gerstaecker

Belonogaster lateritius Gerstaecker, 1857, Monatsber. K. Preuss. Akad. Wiss. Berlin 1857: 463, female -- gMossambiqueh (Berlin); 1862, in Peters, Naturwiss. Reise Mossambique, Hym.: 468, pl. 30 fig. 7. -- Kohl, 1893 (1892), Jahrb. Hamburg. Wiss. Anst. 10 (2): 186; 1894, Ann. Naturhist. Hofmus. Wien 9: 336. -- Gribodo, 1894, in Emery et al., Mem. R. Accad. Sci. Bologna (5) 4: 123. -- Dalla Torre, 1894, Cat. Hym. 9: 116 (lateritia; cat.). -- Gribodo, 1895, Mem. R. Accad. Sci. Bologna (5) 5: 335. -- Dalla Torre, 1904, Genera Insectorum 19: 80 (lateritia; cat.). -- du Buysson, 1908, Résultats Scientif. in Voy. Afrique dfEd. Foá: 593; 1909, Ann. Soc. Entomol. France 78 (2): 216, 218, 256, 264 (male; ethology). -- Bequaert, 1918, Bull. Am. Mus. Nat. Hist. 39: 235 (key), 238, 334 (cat.). -- Schouteden, 1919, Rev. Zool. Afr. 6: 186 (distr.). -- von Schulthess, 1922, Dtsch. Entomol. Z.: 400 (distr.); 1923, in Michaelsen, Beitr. Kennt. Land- und Süsswasserfauna Deutsch-Südwestafrikas 2 (2): 135 (distr.). -- Bequaert, 1938, Explor. Parcs Nation. Congo Belge Miss. G. F. de Witte (1933--1935) 11: 9 (distr.). -- Guiglia, 1940, Atti Mus. Civ. Stor. Nat. Trieste 14 (19): 288. -- Giordani Soika, 1961, S. Afr. Anim. Life 8: 450 (distr.). -- Richards, 1982, Bull. Br. Mus. (Nat. Hist.) Entomol. 44: 36, 37, 44 (key), 72, figs. 32, 38 (lateritia; syn.: B. elegans Gerstaecker, B. fleckii Kohl, B. agilis Kohl; distr.). -- Jeanne et al., 1983, Zoomorphologie 103: 155 (lateritia; morphology).

Belonogaster elegans Gerstaecker, 1857, Monatsb. K. Akad. Wiss. Berlin: 463, female -- gMossambiqueh (type depository unknown); 1862, in Peters, Naturwiss. Reise Mossambique, Hym.: 469, pl. 30 fig. 8. -- Kohl, 1893 (1892), Jahrb. Hamburg. Wiss. Anst. 10 (2): 187; 1894, Ann. Naturhist. Hofmus. Wien 9: 321, 323 (key), 335, pl. 15 fig. 76, pl. 17 figs. 128, 131 (male). -- Gribodo, 1894, in Emery et al., Mem. R. Accad. Sci. Bologna (5) 4: 123. -- Dalla Torre, 1894, Cat. Hym. 9: 114 (cat.). -- Gribodo, 1895, Mem. R. Accad. Sci. Bologna (5) 5: 334. -- Stadelmann, 1898, Dtsch. Ost-Afrika 4, Hym.: 35. -- von Schulthess, 1899, Bull. Soc. Vaud. Sci. Nat. (4), 35: 269 (distr.). -- Dalla Torre, 1904, Genera Insectorum 19: 80 (cat.). -- Zavattari, 1909, Il Ruwenzori, Parte Scientif. 1, Zool.: 211, 271. -- Richards, 1982, Bull. Br. Mus. (Nat. Hist.) Entomol. 44: 72 (syn. of B. lateritia Gerstaecker).

Belonogaster agilis Kohl, 1893 (1892), Jahrb. Hamburg. Wiss. Anst. 10 (2): 187, figs. 1, 4, 9, 10, 15, female, male -- gOst-Afrika: Mbusini (Usegua)h [Angola] (lectotype male Wien); 1894, Ann. Naturhist. Hofmus. Wien 9: 321, 323 (key), 327, 334, pl. 15 fig. 73, pl. 16 figs. 96, 115, pl. 17 figs. 134, 151, 155. -- Dalla Torre, 1894, Cat. Hym. 9: 114 (cat.); 1904, Genera Insectorum 19: 80 (cat.). -- Richards, 1982, Bull. Br. Mus. (Nat. Hist.) Entomol. 44: 72 (designation of lectotype; syn. of B. lateritia Gerstaecker).

Belonogaster fleckii Kohl, 1894 (1893), Ann. Naturhist. Hofmus. Wien 9: 322 (key), 332, 335, female -- gSüdwestafrika: Damaralandh (lectotype Wien). -- Dalla Torre, 1894, Cat. Hym. 9: 115 (cat); 1904, Genera Insectorum 19: 80 (flecki [!]; cat.). -- Bequaert, 1918, Bull. Am. Mus. Nat. Hist. 39: 334 (syn. of B. lateritius var. agilis Kohl). -- Richards, 1982, Bull. Br. Mus. (Nat. Hist.) Entomol. 44: 72 (designation of lectotype; syn. of B. lateritia Gerstaecker).

Belonogaster lateritius var. agilis; du Buysson, 1909, Ann. Soc. Entomol. France 78 (2): 257. -- Bequaert, 1918, Bull. Am. Mus. Nat. Hist. 39: 334 (cat.; syn.: B. fleckii Kohl).

Belonogaster lateritius var. elegans; du Buysson, 1909, Ann. Soc. Entomol. France 78 (2): 257, 264. -- Bequaert, 1918, Bull. Am. Mus. Nat. Hist. 39: 334 (cat.). -- Salt and Bequaert, 1929, Psyche 36: 262 (record of stylopized specimen). -- Giordani Soika, 1935, Mem. Estud. Mus. Zool. Univ. Coimbra (1) 82: 14 (distr.).

Distribution: Somalia; Kenya; Tanzania; Malawi; Angola; Zambia; Zimbabwe; Mozambique; Namibia; Botswana; South Africa (Western Cape, Free State, Natal, Transvaal); Lesotho.

 

leonhardii du Buysson

Belonogaster leonhardii du Buysson, 1909, Ann. Soc. Entomol. France 78 (2): 217, 252, female -- gOuganda anglais: Iraouerh (lectotype Paris). -- Bequaert, 1918, Bull. Am. Mus. Nat. Hist. 39: 335 (cat.); 1938, Explor. Parcs Nation. Congo Belge Miss. G. F. de Witte (1933--1935) 11: 9 (distr.). -- Benoit, 1956, Ann. Mus. R. Congo Belge 8 (Sci. Zool.) 51: 553 (leonhardi [!]). -- Richards, 1982, Bull. Br. Mus. (Nat. Hist.) Entomol. 44: 36, 43 (key), 87 (designation of lectotype), figs. 60--61 (distr.).

Distribution: Uganda; Burundi; Zaire.

 

leonina Richards

Belonogaster leonina Richards, 1982, Bull. Br. Mus. (Nat. Hist.) Entomol. 44: 39, 43 (key), 90, figs. 63--65, male, female -- gSierra Leone: Mattruh (holotype male London); also from Congo; Gabon; Uganda.

Distribution: Sierra Leone; Gabon; Congo; Uganda.

 

levior Richards

Belonogaster levior Richards, 1982, Bull. Br. Mus. (Nat. Hist.) Entomol. 44: 34, 41 (key), 58, female, male -- gUganda: Budongo Forest, Unyoro, 3400 ft [1040 m]h (holotype female London); also from Congo; Liberia.

Distribution: Liberia; Congo; Uganda.

 

libera Richards

Belonogaster libera Richards, 1982, Bull. Br. Mus. (Nat. Hist.) Entomol. 44: 42 (key), 86, male -- gLiberia: Robertsporth (Leiden). -- Kojima and van Achterberg, 1997, Zool. Meded., Leiden 71: 162 (holotype depository).

Distribution: Liberia.

 

longitarsus Richards

Belonogaster longitarsus Richards, 1982, Bull. Br. Mus. (Nat. Hist.) Entomol. 44: 40, 45 (key), 97, figs. 74--76, male, female -- gUganda: banks of Victoria Nile near Masindi Port, 3400 ft [1040m]h (holotype male London); also from three other localities in Uganda; and Zambia; Kenya; Mozambique; South Africa; Tanzania; Angola.

Distribution: Kenya; Uganda; Tanzania; Angola; Zambia; Mozambique; South Africa (Transvaal, Kwazulu-Natal, Western Cape).

 

macilenta (Fabricius)

Vespa macilenta Fabricius, 1781, Spec. Insect. 1: 468 -- gAfrica aequinoctialih (holotype male London); 1787, Mant. Insect. 1: 293. -- Gmelin, 1790, Syst. Nat. ed. 13, 1 (5): 2754. -- Olivier, 1792, Encycl. Méthod. Insect. 6: 673. -- Fabricius, 1793, Entomol. Syst. 2: 280. -- de Saussure, 1853, Ét. Fam. Vesp. 2: 15 (syn. of Raphigaster rufipennis). -- Dalla Torre, 1894, Cat. Hym. 9: 115 (syn. of B. grisea); 1904, Genera Insectorum 19: 80 (syn. of B. grisea). -- Bequaert, 1918, Bull. Am. Mus. Nat. Hist. 39: 331 (syn. of B. grisea (Fabricius)).

Zethus macilentus; Fabricius, 1804, Syst. Piet.: 283.

Belonogaster macilentus; Smith, 1857, Cat. Hym. Br. Mus. 5: 94 (syn.: Raphigastrer filiformis de Saussure, Zethus macilentus (Fabricius)). -- Gribodo, 1894, in Emery et al., Mem. R. Accad. Sci. Bologna (5) 4: 123. -- Richards, 1982, Bull. Br. Mus. (Nat. Hist.) Entomol. 44: 36, 37 (key), 81, figs. 55--57 (macilenta; syn.: B. pusillus Kohl; distr.).

Belonogaster pusillus Kohl, 1894 (1893), Ann. Naturhist. Hofmus. Wien 9: 320, 323 (key), 325, 337, pl. 15 figs. 75, 77, pl. 16 fig. 116, pl. 17, figs. 137, 140, 143, 148, female, male -- gWestafrika: Sierra Leoneh (?Brussels). -- Dalla Torre, 1894, Cat. Hym. 9: 116 (pusilla; cat.); 1904, Genera Insectorum 19: 80 (pusilla; cat.). -- Tullgren, 1904, Ark. Zool 1: 454 (Belenogaster [!]). -- du Buysson, 1909, Ann. Soc. Entomol. France 78 (2): 215, 218, 239, 264, pl. 4 fig. 2 (ethology); 1914, in Voy. Alluaud et Jeannel Afr. Or., Rés. Sci., Ins. Hym. 3, Vesp.: 161. -- Bequaert, 1918, Bull. Am. Mus. Nat. Hist. 39: 235 (key), 236, 336 (cat.; distr.). -- Salt and Bequaert, 1929, Psyche 36: 263 (record of stylopized specimen). -- Bequaert, 1938, Explor. Parcs Nation. Congo Belge Miss. G. F. de Witte (1933--1935) 11: 9 (distr.). -- Richards, 1982, Bull. Br. Mus. (Nat. Hist.) Entomol. 44: 81 (syn. of macilenta).

Distribution: Sierra Leone; Guinea-Bissau; Liberia; Côte dfIvoire; Ghana; Nigeria.

 

maculata Richards

Belonogaster maculata Richards, 1982, Bull. Br. Mus. (Nat. Hist.) Entomol. 44: 36 (key), 71, female -- gUganda: eastern Mbale district, S. of Mt Elgon, 2700--3000 ft [820--915 m]h (Oxford); also from other locality in Uganda; and Kenya.

Distribution: Uganda; Kenya.

 

madecassa de Saussure

Raphigaster madecassus de Saussure, 1853, Ét. Fam. Vesp. 2: 16, pl. 2 fig. 7, female -- gMadagascarh (Genova).

Belonogaster madecassus; Smith, 1857, Cat. Hym. Br. Mus. 5: 94. -- de Saussure, 1890, in Grandidier, Hist. Madagascar 20, Hym. 1: 89 (key), 98. -- Kohl, 1894 (1893), Ann. Naturhist. Hofmus. Wien 9: 337. -- Gribodo, 1894, in Emery et al., Mem. R. Accad. Sci. Bologna (5) 4: 123. -- Dalla Torre, 1894, Cat. Hym. 9: 116 (madecassa; cat.). -- de Saussure, 1900, Abh. Senckenb. Naturforsch. Ges. 26 (2): 207 (key), 211. -- Dalla Torre, 1904, Genera Insectorum 19: 80 (madecassa; cat.). -- du Buysson, 1909, Ann. Soc. Entomol. France 78 (2): 261. -- Bequaert, 1918, Bull. Am. Mus. Nat. Hist. 39: 335 (cat.). -- Richards, 1982, Bull. Br. Mus. (Nat. Hist.) Entomol. 44: 46 (key), 103, figs. 82, 83 (madecassa; distr.). -- Hensen and Blommers, 1987, Tijdschr. Entomol. 130: 12 (madecassum [!]; key), 23, figs. 1, 3, 21, 22, 45 (madecassa; syn.: keiseri Richards; nest; distr.).

Belonogaster longistylus de Saussure, 1890, in Grandidier, Hist. Madagascar 20, Hym. 1: 89 (key), 97, pl. 17 fig. 13, female -- gMadagascarh (lectotype Genève). -- Kohl, 1894 (1893), Ann. Naturhist. Hofmus. Wien 9: 336. -- Dalla Torre, 1894, Cat. Hym. 9: 116 (longistyla; cat.). -- de Saussure, 1900, Abh. Senckenb. Naturforsch. Ges. 26 (2): 207 (key). -- Dalla Torre, 1904, Genera Insectorum 19: 80 (longistyla; cat.). -- von Schulthess, 1907, Voeltzkow, Reise in Ostafrika 1903--1905, Wiss. Ergebn. 2 (2): 64. -- du Buysson, 1909, Ann. Soc. Entomol. France 78 (2): 214, 231. -- Bequaert, 1918, Bull. Am. Mus. Nat. Hist. 39: 335 (cat.). -- Richards, 1982, Bull. Br. Mus. (Nat. Hist.) Entomol. 44: 103, 108, 113 (longestylus [!]; syn. of B. madecassa de Saussure). -- Hensen and Blommers, 1987, Tijdschr. Entomol. 130: 23 (longestylus [!]; syn. of B. madecassa). — Kojima, 2001a, Nat. hist. Bull. Ibaraki Univ. (5): 19 (notes on lectotype).

Belonogaster keiseri Richards, 1982, Bull. Br. Mus. (Nat. Hist.) Entomol. 44: 46 (key), 104, fig. 84, male -- gMadagascar: Diego‑Suarez, Joffrevilleh (Leiden). -- Hensen and Blommers, 1987, Tijdschr. Entomol. 130: 23 (syn. of B. madecassa de Saussure). -- Kojima and van Achterberg, 1997, Zool. Meded., Leiden 71: 162 (holotype depository).

Distribution: Madagascar; Comoros.

 

malagassa de Saussure

Belonogaster malagassus de Saussure, 1900, Abh. Senckenb. Naturforsch. Ges. 26 (2): 207 (key), 210, female -- gMadagascarh (lectotype Genève). -- Dalla Torre, 1904, Genera Insectorum 19: 80 (malagassa; cat.). -- du Buysson, 1909, Ann. Soc. Entomol. France 78 (2): 214, 230. -- Bequaert, 1918, Bull. Am. Mus. Nat. Hist. 39: 335 (cat.). -- Richards, 1982, Bull. Br. Mus. (Nat. Hist.) Entomol. 44: 110 (?syn. of eumenoides de Saussure). -- Hensen and Blommers, 1987, Tijdschr. Entomol. 130: 16 (designation of lectotype; syn. of bicolor). — Kojima, 2001a, Nat. Hist. Bull. Ibaraki Univ. (5): 12 (treating as valid species), 13 (key), figs. 22, 24, 27.

Belonogaster malagassus var. de Saussure, 1900, Abh. Senckenb. Naturforsch. Ges. 26 (2): 210.

Distribution: Madagascar.

 

mandraka Hensen and Blommers

Belonogaster mandraka Hensen and Blommers, 1987, Tijdschr. Entomol. 130: 14 (key), 23, fig. 23, female -- gRép. Malgache, Manjakandriana, Mandraka, 1200 mh (unique Amsterdam). -- Kojima and van Achterberg, 1997, Zool. Meded., Leiden 71: 167 (holotype depository).

Distribution: Madagascar.

 

maromandia Richards

Belonogaster maromandia Richards, 1982, Bull. Br. Mus. (Nat. Hist.) Entomol. 44: 47 (key), 108, female -- gMadagascar: Province de Amalalava, Maromandiah (Paris). -- Hensen and Blommers, 1987, Tijdschr. Entomol. 130: 14 (key), 24, figs. 40, 41 (male; distr.).

Distribution: Madagascar.

 

menelikii Gribodo

Belonogaster menelikii Gribodo, 1879, Ann. Mus. Civ. Genova (1) 14: 342, female -- gMahal-Uonzh (Genova); 1881, Ann. Mus. Civ. Genova 16: 288 (male); 1884, Ann. Mus. Civ. Genova 20: 384; 1884, Ann. Mus. Civ. Genova 21: 288. -- Kohl, 1894 (1893), Ann. Naturhist. Hofmus. Wien 9: 320, 323 (key), 337, pl. 15 figs. 81, 92, pl. 17 fig. 135. -- Gribodo, 1894, in Emery et al., Mem. R. Accad. Sci. Bologna (5) 4: 123. -- Dalla Torre, 1894, Cat. Hym. 9: 116 (cat.). -- Fox, 1896, Proc. Acad. Nat. Sci. Philadelphia 48: 555. -- Dalla Torre, 1904, Genera Insectorum 19: 80 (meneliki [!]; cat.). -- Schulz, 1911, Zool. Ann. Würzburg 4: 200. -- Richards, 1982, Bull. Br. Mus. (Nat. Hist.) Entomol. 44: 35, 38, 43 (key), 52, 113, fig. 8 (meneliki [!]; distr.).

Belonogaster griseus var. menelikii; du Buysson, 1909, Ann. Soc. Entomol. France 78 (2): 250, 265, pl. 7 fig. 1 (ethology) [partim]; 1914, in Voy. Alluaud et Jeannel Afr. Or. Rés. Sci., Ins. Hym. 3, Vesp.: 163. -- Bequaert, 1918, Bull. Am. Mus. Nat. Hist. 39: 332 (cat.). -- Schouteden, 1919, Rev. Zool. Afr. 6: 186 (distr.). -- von Schulthess, 1928, Senckenbergiana 10 (3/4): 96 (distr.). -- Guiglia, 1932 (1931), Mem. Soc. Entomol. Ital. 10: 119 (meneliki [!]; distr.). -- Bequaert, 1933, Ann. Mag. Nat. Hist. 12: 116 (possibly distinct species; distr.; records by du Buysson, 1909, Schouteden, 1919, von Schulthess, 1928, dubious). -- Guiglia, 1939, Mem. Soc. Entomol. Ital. 17: 192 (meneliki [!]); 1939, Mem. Soc. Entomol. Ital. 18: 88 (meneliki [!]); 1940, Atti Mus. Civ. Stor. Nat. Trieste 14 (19): 288 (meneliki [!]). -- Giordani Soika, 1940, Atti Mus. Civ. Stor. Nat. Trieste 14 (18): 286 (meneliki [!]; distr.); 1952 (1951), Riv. Biol. Colon. 11: 88 (meneliki [!]; distr.). -- Iwata, 1966, Mushi 39: 57 (ethology); 1976, Evol. Instinct: 303, 307, 308 (ethology).

Distribution: Ethiopia.

 

multipunctata Richards

Belonogaster multipunctata Richards, 1982, Bull. Br. Mus. (Nat. Hist.) Entomol. 44: 36 (key), 55, female -- gUganda: Entebbe, 3800--4000 ft [1160--1220 m], forest within 4 miles [6.5 km] of Kitabi Hillh (Oxford).

Distribution: Uganda.

 

neavei Richards

Belonogaster neavei Richards, 1982, Bull. Br. Mus. (Nat. Hist.) Entomol. 44: 39, 45 (key), 89, male, female -- gKenya: Masongaleni, ca 38‹2fE, 2‹4fS, 3000 ft [915 m]h (holotype male London); also from 6 other localities in Kenya; and Zambia; Malawi; Tanzania; South Africa.

Distribution: Kenya; Tanzania; Zambia; Malawi; South Africa (Transvaal).

 

nigricans Richards

Belonogaster nigricans Richards, 1982, Bull. Br. Mus. (Nat. Hist.) Entomol. 44: 36 (key), 77, fig. 48, female -- gCongo: Dimonikah (holotype Paris); also from 3 other localities in Congo.

Distribution: Congo.

 

nitida Richards

Belonogaster nitida Richards, 1982, Bull. Br. Mus. (Nat. Hist.) Entomol. 44: 40 (key), 60, figs. 15, 16, male -- gNigeria: Eastern province, Oguta, east of Onitshah (unique London).

Distribution: Nigeria.

 

ornata de Saussure

Belonogaster ornatus de Saussure, 1900, Abh. Senckenb. Naturforsch. Ges. 26 (2): 207 (key), 209, female, male -- gMadagascarh (lectotype male Genève). -- Dalla Torre, 1904, Genera Insectorum 19: 80 (ornata; cat.). -- Bequaert, 1918, Bull. Am. Mus. Nat. Hist. 39: 330 (syn. of B. eumenoides de Saussure). -- Richards, 1982, Bull. Br. Mus. (Nat. Hist.) Entomol. 44: 110 (?syn. of B. eumenoides de Saussure). -- Hensen and Blommers, 1987, Tijdschr. Entomol. 130: 14 (key), 25, figs. 24, 25 (ornata; designation of lectotype; syn.: pomicolor de Saussure; distr.). — Kojima, 2001a, Nat. hist. Bull. Ibaraki Univ. (5): 16 (ornata), figs. 48-53, 19 (key).

 

pomicolor de Saussure

Belonogaster pomicolor de Saussure, 1900, Abh. Senckenb. Naturforsch. Ges. 26 (2): 207, 209, female, male -- gMadagascarh (lectotype female Genève). -- Dalla Torre, 1904, Genera Insectorum 19: 80 (cat.). -- Bequaert, 1918, Bull. Am. Mus. Nat. Hist. 39: 330 (syn. of B. eumenoides de Saussure). -- Richards, 1982, Bull. Br. Mus. (Nat. Hist.) Entomol. 44: 110 (?syn. of eumenoides de Saussure). -- Hensen and Blommers, 1987, Tijdschr. Entomol. 130: 25, 26 (designation of lectotype; syn. of B. ornata de Saussure). — Kojima, 2001a, Nat. hist. Bull. Ibaraki Univ. (5): 17 (taxonomy; different species from B. ornata; description of male), figs. 61-72, 19 (key).

Distribution: Madagascar.

 

pennata Richards

Belonogaster pennata Richards, 1982, Bull. Br. Mus. (Nat. Hist.) Entomol. 44: 36, 43 (key), 51, fig. 7, female, male -- gZaire: Shaba prov. (Katanga), Kambove, 4000--5000 ft [1220--1500 m]h (holotype female London); also from 6 other localities in Zaire; and Gabon; Congo; Rwanda; Angola; Kenya; Uganda; Tanzania; Zambia; Malawi; Zimbabwe; South Africa.

Distribution: Gabon; Congo; Zaire; Uganda; Kenya; Rwanda; Tanzania; Angola; Zambia; Malawi; Zimbabwe; South Africa.

 

petiolata (DeGeer)

Vespa petiolata DeGeer, 1778, Mém. Hist. Insectes 7: 610, pl. 45, fig. 10 -- gCap-de-Bonne-Esperanceh (holotype female Stockholm). -- Göze, 1783, in DeGeer, Abh. Gesch. Insekten 7: 217, T. 45, F. 10. -- Retzius, 1783, Gen. Spec. Insect. 6: 64.

?Vespa linearis Olivier, 1792, Encycl. Méthod. Ins. 6: 673 -- gCap de Bonne Espéranceh (lost). -- Dalla Torre, 1894, Cat. Hym. 9: 116 (?syn. of B. petiolata). -- Bequaert, 1918, Bull. Am. Mus. Nat. Hist. 39: 331 (syn. of B. grisea (Fabricius)).

?Polistes linearis; Latreille, 1809, Gen. Crust. Insectes 4: 142.

Belonogaster brachycerus Kohl, 1894 (1893), Ann. Naturhist. Hofmus. Wien 9: 323, 324, 331, pl. 15 fig. 82, pl. 17 fig. 138, female -- gSüdafrika: Cap b. sp.h (type depository unknown). -- Dalla Torre, 1894, Cat. Hym. 9: 114 (brachycera; cat.); 1904, Genera Insectorum 19: 80 (brachycera; cat.). -- du Buysson, 1909, Ann. Soc. Entomol. France 78 (2): 217, 218, 254, 264 (male; ethology). -- von Schulthess, 1914, Dtsch. Entomol. Z. 1914: 290 (compared to B. rothkirchi n. sp.). -- Bequaert, 1918, Bull. Am. Mus. Nat. Hist. 39: 335 (syn. of B. petiolata (DeGeer)).

?Belonogaster linearis; Kohl, 1894 (1893), Ann. Naturhist. Hofmus. Wien 9: 336. -- Dalla Torre, 1894, Cat. Hym. 9: 116 (cat.); 1904, Genera Insectorum 19: 80 (cat.).

Belonogaster petiolatus; Kohl, 1894 (1893), Ann. Naturhist. Hofmus. Wien 9: 337. -- Dalla Torre, 1894, Cat. Hym. 9: 116 (petiolata; cat.; error: gPolistes petiolata Lepeletier, Encycl. méthod. Insect. X. 1825 p. 173 n. 16h [absent in Lepeletier, 1825]); 1904, Genera Insectorum 19: 80 (petiolata; cat.). -- Tullgren, 1904, Ark. Zool. 1: 461 (Belenogaster [!]). -- Schulz, 1912, Berlin. Entomol. Z. 57: 62. -- Bequaert, 1918, Bull. Am. Mus. Nat. Hist. 39: 335 (cat.; syn.: B. brachycerus Kohl). -- Richards, 1982, Bull. Br. Mus. (Nat. Hist.) Entomol. 44: 35, 37, 43 (key), 71, figs. 36, 37 (petiolata; syn.: B. brachycerus Kohl, ?V. linearis Olivier; distr.). -- Keeping and Crewe, 1983, J. Entomol. Soc. South. Africa 46: 309 (petiolata; ethology.). -- Kojima and Keeping, 1984, J. Entomol. Soc. South. Africa 48: 237 (petiolata; larva). -- Keeping et al., 1986, J. Chem. Ecol. 773 (petiolata; ethology). -- Keeping and Crewe, 1987, in Eder and Rembold, Chem. Biol. Soc. Insects: 383 (petiolata; ethology). -- Keeping, 1990, J. Insect Behav. 3 (1): 85 (petiolata; ethology); 1990, Ethology 85: 1  (petiolata; ethology); 1990, J. Entomol. Soc. South. Africa 54: 17 (petiolata; ethology). -- Spradbery, 1991, in Ross and Matthews, Soc. Biol. Wasps: 354, 355, 357, 373 (petiolata; colony population). -- Jeanne, 1991, in Ross and Matthews, Soc. Biol. Wasps: 393 (petiolata; ethology). -- Ross and Carpenter, 1991, in Ross and Matthews, Soc. Biol. Wasps: 464 (petiolata; biology). -- Keeping, 1992, Behav. Ecol. Sociobiol. 31: 211 (petiolata; ethology). -- Itô, 1993, Behav. Soc. Evol. Wasps: 20, 34, 104 (petiolata; ethology). -- Keeping, 1995, Insectes Soc. 42 (3): 317 (petiolata; ethology).

Distribution: Malawi; Zimbabwe; South Africa (Western Cape, Free State, Transvaal, Kwazulu-Natal); Lesotho.

 

pileata Richards

Belonogaster pileata Richards, 1982, Bull. Br. Mus. (Nat. Hist.) Entomol. 44: 37, 41 (key), 62, figs. 21, 22, female, male -- gKenya: Tareta forest (near Tanzanian border)h (holotype female London); also from three other localities in Kenya; and Tanzania; Malawi.

Distribution: Kenya; Tanzania; Malawi.

 

prasina de Saussure

Belonogaster prasinus de Saussure, 1890, in Grandidier, Hist. Madagascar 20, Hym. 1: 89 (key), 92, pl. 19 fig. 5, female, male -- gMadagascar ... aux environs de Fort-Dauphinh (lectotype male Genève); also from 2 other localties. -- Kohl, 1894 (1893), Ann. Naturhist. Hofmus. Wien 9: 337. -- Dalla Torre, 1894, Cat. Hym. 9: 116 (prasina; cat.). -- de Saussure, 1900, Abh. Senckenb. Naturforsch. Ges. 26 (2): 207 (key). -- Dalla Torre, 1904, Genera Insectorum 19: 80 (prasina; cat.). -- von Schulthess, 1907, Voeltzkow, Reise in Ostafrika 1903--1905, Wiss. Ergebn. 2 (2): 65. -- du Buysson, 1909, Ann. Soc. Entomol. France 78 (2): 214, 228. -- Bequaert, 1918, Bull. Am. Mus. Nat. Hist. 39: 335 (cat.). --  Richards, 1982, Bull. Br. Mus. (Nat. Hist.) Entomol. 44: 47 (key), 107, figs. 87, 89 (prasina; designation of lectotype; distr.). -- Hensen and Blommers, 1987, Tijdschr. Entomol. 130: 14 (key), 26 (prasina; distr.). — Kojima, 2001a, Nat. Hist. Bull. Ibaraki Univ. (5): 19 (prasina; notes on lectotype and paralectotypes).

Distribution: Madagascar.

 

principalis Richards

Belonogaster principalis Richards, 1982, Bull. Br. Mus. (Nat. Hist.) Entomol. 44: 38 (key), 82, female -- gGuinea-Bissau: Principé I., between Roca Esperanza and Roca Sundih [sic] (unique London).

Distribution: Principé.

 

punctata Richards

Belonogaster punctata Richards, 1982, Bull. Br. Mus. (Nat. Hist.) Entomol. 44: 42 (key), 84, male -- gCameroun: Kumbah (unique London).

Distribution: Cameroon.

 

punctilla Richards

Belonogaster punctilla Richards, 1982, Bull. Br. Mus. (Nat. Hist.) Entomol. 44: 38 (key), 83, female -- gUganda: Mpanga forest, Toro, 4800ft [1460m]h (unique London).

Distribution: Uganda.

 

pusilloides Richards

Belonogaster pusilloides Richards, 1982, Bull. Br. Mus. (Nat. Hist.) Entomol. 44: 37, 43 (key), 80, fig. 54, female, male -- gUganda: top of escarpment, east of Batiaba, L. Albert, 3200 ft [980m]h (holotype female London); also from two other localities in Uganda; and Kenya; Congo; South Africa. — Kojima, 2001a, Nat. Hist. Bull. Ibaraki Univ. (5): 20.

Distribution: Congo; Uganda; Kenya; South Africa (Kwazulu-Natal).

 

rothkirchi von Schulthess

Belonogaster rothkirchi von Schulthess, 1914, Soc. Entomol. 29: 4, male -- gKamerunberg, Soppoh (lectotype Zürich); 1914, Dtsch. Entomol. Z.: 288 (fig.). -- Bequaert, 1918, Bull. Am. Mus. Nat. Hist. 39: 336 (cat.). -- Richards, 1982, Bull. Br. Mus. (Nat. Hist.) Entomol. 44: 42, 45 (key), 85, fig. 59 (designation of lectotype).

Distribution: Cameroon.

 

saeva de Saussure

Belonogaster saevus de Saussure, 1890, in Grandidier, Hist. Madagascar 20, Hym. 1: 92, footnote, female -- glfAfrique tropicaleh (type depository unknown). -- Kohl, 1894 (1893), Ann. Naturhist. Hofmus. Wien 9: 337. -- Dalla Torre, 1894, Cat. Hym. 9: 116 (saeva; cat.); 1904, Genera Insectorum 19: 80 (saeva; cat.). -- du Buysson, 1909, Ann. Soc. Entomol. France 78 (2): 215, 219, 242, 264, pl. 4 figs. 10, 11 (male). -- von Schulthess, 1912, Wiss. Ergebn. D. Z. Afrik. Exp., (1907--08), 4, Lf. 10: 291; 1913, Mitt. Zool. Mus. Berlin 6 (3): 338. -- Bequaert, 1918, Bull. Am. Mus. Nat. Hist. 39: 234 (key), 336 (cat.; distr.). -- Schouteden, 1919, Rev. Zool. Afr. 6: 184 (distr.). -- Bequaert, 1938, Explor. Parcs Nation. Congo Belge Miss. G. F. de Witte (1933--1935) 11: 9 (distr.). -- Giordani Soika, 1977, Steenstrupia 4: 127 (distr.). -- Richards, 1982, Bull. Br. Mus. (Nat. Hist.) Entomol. 44: 38, 48 (key), 61, figs. 19, 20 (saeva; distr.). — Kojima, 2001a, Nat. Hist. Bull. Ibaraki Univ. (5): 20 (saeva; taxonomic notes).

Distribution: Sierra Leone; Ghana; Cameroon; Congo; Zaire; Uganda; Kenya; Tanzania; Malawi; South Africa (Cape of Good Hope).

 

saussurei Kirby

Belonogaster saussurei Kirby, 1881, Proc. Zool. Soc. London: 649, female, male -- gSocotrah (lectotype male London). -- Kohl, 1894 (1893), Ann. Naturhist. Hofmus. Wien 9: 337. -- Dalla Torre, 1894, Cat. Hym. 9: 116 (saussureii [!]; cat.). -- Kirby, 1898, Proc. Zool. Soc. London: 386; 1903, in Forbes, Nat. Hist. of Sokotra, Zool. Hym.: 248, pl. 16 fig. 2 (Belenogaster [!]; syn.: B. tricolor Taschenberg; ethology). -- Dalla Torre, 1904, Genera Insectorum 19: 80 (cat.). -- Kohl, 1907, Denkschr. K. Akad. Wiss. Wien, Math. Naturh. Kl. 71 (1): 223, pl. 5 figs. 18, 20, 31. -- du Buysson, 1909, Ann. Soc. Entomol. France 78 (2): 262. -- Bequaert, 1918, Bull. Am. Mus. Nat. Hist. 39: 336 (cat.; syn.: B. tricolor Taschenberg). -- Richards, 1982, Bull. Br. Mus. (Nat. Hist.) Entomol. 44: 40, 44 (key), 93, fig. 69 (designation of lectotype; syn.: B. tricolor Taschenberg; distr.); 1985 (1984), Fauna Saudi Arabia 6: 436, 437 (key; distr.), 438 (list).

Belonogaster tricolor Taschenberg, 1883, Z. Ges. Naturw. Halle 56: 175, female -- gInsel Socotra (Socotora)h (type depository unknown). -- Kohl, 1894 (1893), Ann. Naturhist. Hofmus. Wien 9: 337. -- Dalla Torre, 1894, Cat. Hym. 9: 116 (cat.). -- Kirby, 1903, in Forbes, Nat. Hist. of Sokotra, Zool. Hym.: 248 (Belenogaster [!]; syn. of B. saussurei). -- Dalla Torre, 1904, Genera Insectorum 19: 80 (cat.). -- du Buysson, 1909, Ann. Soc. Entomol. France 78 (2): 261. -- Bequaert, 1918, Bull. Am. Mus. Nat. Hist. 39: 336 (syn. of B. saussurei Kirby).

Distribution: Yemen (including Socotra).

 

schulthessi Kojima

Belonoaster schulthessi Kojima, 2001a, Nat. Hist. Bull. Ibaraki Univ. (5): 14, figs. 36-47, male, female – "Rogez, O. Madagascar" (holotype male Zürich), 19 (key).

Distribution: Madagascar.

 

scutifera Hensen and Blommers

Belonogaster scutifera Hensen and Blommers, 1987, Tijdschr. Entomol. 130: 12 (key), 26, figs. 26--28, male, female -- gMadagascar Est: Ambodivoangyh (holotype male Tervuren); also from other locality in Madagascar.

Distribution: Madagascar.

 

somereni Richards

Belonogaster somereni Richards, 1982, Bull. Br. Mus. (Nat. Hist.) Entomol. 44: 35, 44 (key), 50, fig. 6, male, female -- gKenya: Teita Hills (ca 38‹3fE, 2‹4fS)h (holotype male London); also from Tanzania; Zimbabwe; Zambia; Malawi; Mozambique; South Africa; Niger.

Distribution: Niger; Kenya; Tanzania; Zambia; Malawi; Zimbabwe; Mozambique; South Africa (Free State, Kwazulu-Natal).

 

tanosy Hensen and Blommers

Belonogaster tanosy Hensen and Blommers, 1987, Tijdschr. Entomol. 130: 14 (key), 27, figs. 29--31, male, female -- gMadagascar, Fort Dauphin, 500 mh (holotype male London); also from four other localities in Madagascar.

Distribution: Madagascar.

 

tarsata Kohl

Belonogaster tarsatus Kohl, 1893 (1892), Jahrb. Hamburg. Wiss. Anst. 10 (2): 187, figs. 2, 5, 7, 8, 11, 16, female, male -- gOst-Afrika: Mbusini (Usegua ...)h (lectotype male Wien); 1894, Ann. Naturhist. Hofmus. Wien 9: 321, 323 (key), 333, 337, pl. 15 figs. 85, 86, pl. 16 figs. 97, 119, pl. 17 figs. 150, 153, 154. -- Dalla Torre, 1894, Cat. Hym. 9: 116 (tarsata; cat.). -- Stadelmann, 1898, Deutsch Ost-Afrika 4, Hym.: 35. -- Dalla Torre, 1904, Genera Insectorum 19: 80 (tarsata; cat.). -- du Buysson, 1909, Ann. Soc. Entomol. France 78 (2): 215, 219, 237, 263, pl. 4 fig. 7. -- Bequaert, 1918, Bull. Am. Mus. Nat. Hist. 39: 336 (cat.). -- Richards, 1982, Bull. Br. Mus. (Nat. Hist.) Entomol. 44: 37, 44 (key), 73, figs. 39--41 (tarsata; designation of lectotype; distr.).

Distribution: Tanzania (including Zanzibar).

 

tessmanni von Schulthess

Belenogaster [!] tessmanni von Schulthess, 1910, Soc. Entomol. 25: 45, female, male -- gSpanisch Guinea ... Uelleborg, ... Benitogebiet ... Alcu Benitogebiet ... Spanisch Guinea Hinterland, Nkolontanganh (Berlin); 1913, Mitt. Zool. Mus. Berlin 6 (3): 339. -- Bequaert, 1918, Bull. Am. Mus. Nat. Hist. 39: 337 (Belonogaster; cat.); 1938, Explor. Parcs Nation. Congo Belge Miss. G. F. de Witte (1933--1935) 11: 9 (Belonogaster; ?Zaire). -- Richards, 1982, Bull. Br. Mus. (Nat. Hist.) Entomol. 44: 101 (Belonogaster; types not found; gI have not been able to identify this species with any certainty, though it is probably distincth) [error: type locality Guinea-Bissau].

Distribution: Equatorial Guinea; Cameroon; ?Zaire.

 

tipuliformis Hensen and Blommers

Belonogaster tipuliformis Hensen and Blommers, 1987, Tijdschr. Entomol. 130: 12 (key), 28, fig. 32, female -- gMadagascar; (Tamat.); Périnet; 950 m; 48‹16E, 18‹56Sh (holotype Leiden); also from two other localities in Madagascar. -- Kojima and van Achterberg, 1997, Zool. Meded., Leiden 71: 162 (holotype depository).

Distribution: Madagascar.

 

trandraka Hensen and Blommers

Belonogaster trandraka Hensen and Blommers, 1987, Tijdschr. Entomol. 130: 12 (key), 30, figs. 33, 34, female -- gMadagascar Est: Ambodivoangyh (Tervuren).

Distribution: Madagascar.

 

turbulenta Kohl

Belonogaster turbulentus Kohl, 1894 (1893), Ann. Naturhist. Hofmus. Wien 9: 323 (key), 330, 337, pl. 15 fig. 87, pl. 16 fig. 101, pl. 17 fig. 147, female -- gWestafrika: Sierra Leoneh (Wien). -- Dalla Torre, 1894, Cat. Hym. 9: 116 (turbulenta; cat.); 1904, Genera Insectorum 19: 80 (turbulenta; cat.). -- Tullgren, 1904, Ark. Zool. 1: 454 (Belenogaster [!]). -- du Buysson, 1909, Ann. Soc. Entomol. France 78 (2): 215, 216, 240, 264, pl. 2 fig. 13. -- von Schulthess, 1913, Mitt. Zool. Mus. Berlin 6 (3): 338; 1914, Dtsch. Entomol. Z. 1914: 290 (compared to B. rothkirchi n. sp.) -- Bequaert, 1918, Bull. Am. Mus. Nat. Hist. 39: 235 (key), 236, 337 (cat.; distr.). -- Schouteden, 1919, Rev. Zool. Afr. 6: 185 (distr.). -- von Schulthess, 1928, Senckenbergiana 10 (3/4): 96 (distr.). -- Salt and Bequaert, 1929, Psyche 36: 263 (record of stylopized specimen). -- Bequaert, 1938, Explor. Parcs Nation. Congo Belge Miss. G. F. de Witte (1933--1935) 11: 8, 9, 10 (distr.). -- Richards, 1982, Bull. Br. Mus. (Nat. Hist.) Entomol. 44: 36 (key), 74, fig. 42 (turbulenta; distr.).

Distribution: Sierra Leone; Congo; Zaire.

 

turgida Kohl

Belonogaster turgidus Kohl, 1894 (1893), Ann. Naturhist. Hofmus. Wien 9: 322 (key), 333, 337, pl. 15 fig. 74, pl. 16 fig. 114, female -- gFernando Poh (Wien). -- Dalla Torre, 1894, Cat. Hym. 9: 116 (turgida; cat.); 1904, Genera Insectorum 19: 80 (turgida; cat.). -- du Buysson, 1909, Ann. Soc. Entomol. France 78 (2): 260. -- Bequaert, 1918, Bull. Am. Mus. Nat. Hist. 39: 337 (cat.). -- Richards, 1982, Bull. Br. Mus. (Nat. Hist.) Entomol. 44: 34 (key), 60, fig. 17 (turgida).

Distribution: Bioko.

 

ugandae Richards

Belonogaster ugandae Richards, 1982, Bull. Br. Mus. (Nat. Hist.) Entomol. 44: 42 (key), 54, fig. 12, male -- gUganda: eastern Mbale district, S. of Mt Elgon, 3700--3900 ft [1130--1190m]h (London).

Distribution: Uganda.

 

vadoni Hensen and Blommers

Belonogaster vadoni Hensen and Blommers, 1987, Tijdschr. Entomol. 130: 14 (key), 30, female -- gN. E. Madagascar, Fampanamboh (Tervuren).

Distribution: Madagascar.

 

vasseae du Buysson

Belonogaster vasseae du Buysson, 1906, Bull. Soc. Entomol. France 1906: 189, female -- gbassin inférieur du Zambèze, de la vallée de la Muza, à une altitude de 1.000 à 1.200 mètresh (lectotype Paris); 1909, Ann. Soc. Entomol. France 78 (2): 213, 219, 263, pl. 2 fig. 8 (ethology). -- von Schulthess, 1912, Soc. Entomol. 27: 42; 1913, Mitt. Zool. Mus. Berlin 6 (3): 337. -- du Buysson, 1914, in Voy. Alluaud et Jeannel, Afr. Or., Rés. Sci., Ins. Hym. 3, Vesp.: 160 (male). -- Bequaert, 1918, Bull. Am. Mus. Nat. Hist. 39: 234, 337 (cat.). -- Schouteden, 1919, Rev. Zool. Afr. 6: 184 (distr.). -- Bequaert, 1938, Explor. Parcs Nation. Congo Belge Miss. G. F. de Witte (1933--1935) 11: 9 (distr.). -- Benoit, 1956, Ann. Mus. R. Congo Belge 8 (Sci. Zool.) 51: 552 (distr.). -- Giordani Soika, 1977, Steenstrupia 4: 127 (distr.). -- Richards, 1982, Bull. Br. Mus. (Nat. Hist.) Entomol. 44: 35, 41 (key), 56, fig. 13 (designation of lectotype; distr.).

Distribution: Cameroon; Congo; Zaire; Uganda; Rwanda; Kenya; Tanzania; Angola; Zambia; Malawi; Zimbabwe; Mozambique.

 

Species originally described as Belonogaster, now placed in another genus

 

Belenogaster [!] bidentatus Kirby, 1884, Ann. Mag. Nat. Hist. (5) 13: 410, female [male, according to Bequaert, 1928] -- gPandana, Fiji, Aug. 1874h (London). -- Dalla Torre, 1894, Cat. Hym. 9: 114 (bidentata; cat.) = Eumenes [Delta] insularis Smith, 1857, syn. by Bequaert, 1928, Ann. Mag. Nat. Hist. (10) 2: 164.

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