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DISTRIBUTIONAL CHECKLIST OF THE SPECIES OF THE GENUS POLISTES (HYMENOPTERA: VESPIDAE; POLISTINAE, POLISTINI)

 

JAMES M. CARPENTER

Curator, Department of Entomology, American Museum of Natural History.

 

TRIBE POLISTINI LEPELETIER, 1836

 

GENUS POLISTES LATREILLE

 

SUBGENUS GYROSTOMA KIRBY

 

Gyrostoma Kirby, 1828, in Kirby and Spence, Introd. Entomol. Ed. 5 (of whole work), Ed. 2 (of vol. 3): 36, 631, replacement name for Cyclostoma Kirby, 1826, non Lamarck, 1799.

Cyclostoma Kirby, 1826, in Kirby and Spence, Introd. Entomol. Ed. 1, 3: 36, 633, genus. Non Cyclostoma Lamarck, 1799 (Mollusca).

Type species: Cyclostoma gigas Kirby, 1826, by monotypy.

Megapolistes van der van der Vecht, 1968, Bijdr. Dierk. 38: 97, subgenus of Polistes Latreille.

Type species: Vespa olivacea DeGeer, 1773, by original designation.

Nygmopolistes Richards, 1973, Rev. Bras. Entomol. 17: 91, 93, 98, subgenus of Polistes Latreille.

Type species: gPolistes sulcatus Smith, 1852 (= P. rugifrons Cameron, 1900),h by original designation.

Megapostes [!]; Lee, 1982, Hornets Agric. Regions China: 122, 124, 126.

 

arthuri Cameron

Polistes arthuri Cameron, 1901, Proc. Zool. Soc. London 1901: 241.

Polistes tepidus novae-pomeraniae Schulz, 1905, Berlin Entomol. Zeitschr. 49: 227.

Polistes novae pomerania [!]; Cheesman, 1951, Ann. Mag. Nat. Hist. (12)4: 985.

Polistes tepidus novae-pomeranieae [!]; Yoshikawa, 1962, J. Biol. Osaka City Univ. 13: 39.

Polistes novopomeraniae [!]; Richards, 1973, Rev. Bras. Entomol. 17: 100.

Polistes arthuri vechti Petersen, 1990, Entomol. Scand. 21: 65, non Das and Gupta, 1989.

Distribution: Bismarck Archipelago: New Britain, New Ireland (Petersen, 1990, Entomol. Scand. 21).

 

balder Kirby

Polistes balder Kirby, 1888, Proc. Zool. Soc. London 1888: 552.

Polistes bader [!]; Richards, 1978, Aust. J. Zool., Suppl. Ser., 61: 1.

Distribution: Australia: Christmas Is., Cocos Is. (Richards, 1978, Aust. J. Zool., Suppl. Ser., 61).

 

diabolicus de Saussure

Polistes diabolicus de Saussure, 1853, Ét. Fam. Vesp. 2: 68.

Distribution: Indonesia: Java, Lombok, Sumbawa, Flores, Timor, Sumba, (?) Kai Is. (Saito et al., 2005, J. Hym. Res. 140. - Kojima et al., 2006, Serangga 11 (1-2): 11).

 

erythrinus Holmgren

Polistes erythrinus Holmgren, 1868, K. Svenska Fregat. Eugenies Resa Omkring Jorden, Zool. 1, Insect., Hym.: 440.

Distribution: Australia: Queensland, New South Wales, A. C. T., Victoria, South Australia (Richards, 1978, Aust. J. Zool., Suppl. Ser., 61).

 

facilis de Saussure

Polistes facilis de Saussure, 1853, Ét. Fam. Vesp. 2: 53.

Distribution: Australia: Queensland (Richards, 1978, Aust. J. Zool., Suppl. Ser., 61).

 

fordi van der Vecht

Polistes fordi van der Vecht, 1972 (1971), Entomol. Essays to commemorate the retirement of Prof. K. Yasumatsu: 92.

Distribution: Solomon Is.: Bougainville.

 

gigas (Kirby)

Cyclostoma gigas Kirby, 1826, in Kirby and Spence, Introd. Entomol. Ed. 1, 3: 36.

Polistes orientalis Lepeletier, 1836, Hist. Nat. Insect. Hym. 1: 519.

Polistes gyrostoma de Saussure, 1854, Ét. Fam. Vesp. 2: 104.

Gyrostoma orientalis gKirbyh de Saussure, 1854, Ét. Fam. Vesp. 2: 104, as synonym of Polistes gyrostoma de Saussure.  Unavailable under Art. 11.6 of the International Code of Zoological Nomenclature.

Polistes confusus Smith, 1857, Cat. Hym. Br. Mus. 5, Vesp.: 102.

Distribution: India: Sikkim; southern China; Taiwan (Das and Gupta, 1989, Orient. Insect. Monogr. 11).

 

hebridensis erromangensis Giordani Soika

Polistes hebridensis erromangensis Giordani Soika, 1981 (1980), Boll. Mus. Civ. Venezia 31: 118.

Distribution: Vanuatu: Erromango.

 

hebridensis hebridensis Giordani Soika

Polistes hebridensis Giordani Soika, 1981 (1980), Boll. Mus. Civ. Venezia 31: 117.

Distribution: Vanuatu: Anatom.

 

hebridensis malekulensis Giordani Soika

Polistes hebridensis malekulensis Giordani Soika, 1981 (1980), Boll. Mus. Civ. Venezia 31: 119.

Distribution: Vanuatu: Malakula.

 

hebridensis vilensis Giordani Soika

Polistes hebridensis vilensis Giordani Soika, 1981 (1980), Boll. Mus. Civ. Venezia 31: 119.

Distribution: Vanuatu: Port-Vila.

 

jokahamae Radoszkowski

Polistes jokahamae Radoszkowski, 1887, Horae Entomol. Soc. Ross. 21: 435.

Polistes japonicus Cameron, 1900, Ann. Mag. Nat. Hist. (7) 6: 417, non de Saussure, 1858.

Polistes jocahamae [!]; Dalla Torre, 1904, Genera Insect. 19: 70.

Polistes jadwigae Dalla Torre, 1904, Genera Insect. 19: 70.  Replacement name for japonicus Cameron, 1900, non de Saussure, 1858.

Polistes perkinsi Kohl, 1908, Denkschr. Akad. Wiss., Wien 81: 313.

Polistes yokohamae; Matsumura, 1911, Thousand Insect. Japan, Suppl. 3: 105, unnecessary emendation.

Polistes okinawensis Matsumura and Uchida, 1926, Insect. Matsum. 1 (1): 34.

Polistes macaensis tahitensis Cheesman, 1928, Ann. Mag. Nat. Hist. (10) 1: 180, male, female - gTuamotu Arch.: Napuka, ... Society Is.: Tahiti, ... Bora Borah (type not indicated, London.

Polistes perkinsii [!]; Liu, 1937, Peking Nat. Hist. Bull. 11 (3): 313.

Polistes fadwigae [!]; Yoshikawa, 1962, J. Biol. Osaka City Univ. 13: 20.

Polistes jokohamae Yasumatsu, 1940, Trans. Sapporo Nat. Hist. Soc. 16: 96, unnecessary emendation.

Polistes yokahamae [!]; Sonan, 1943, Trans. Nat. Hist. Soc. Taiwan 33 (242-243): 475.

Polistes okinawaensis [!]; Yoshikawa, 1962, J. Biol. Osaka City Univ. 13: 33.

Distribution: India; China; Mongolia; Korea; Japan, including Ryukyu Is.; Taiwan; Society Islands: Bora Bora, Tahiti; Tuamotu Archipelago: Napuka; introduced U.S.A.: Hawaii (Richards, 1978, Soc. Wasps Am.; Kim, 1980, Distr. Atlas Insect. Korea, Ser. 3, Hym. Dipt.; Starr, 1992, Bull. Natl. Mus. Nat. Sci. 3).

 

legnotus van der Vecht

Polistes legnotus van der Vecht, 1972 (1971), Entomol. Essays to commemorate the retirement of Prof. K. Yasumatsu: 100.

Distribution: Solomon Is.: Guadalcanal.

 

olivaceus (DeGeer)

Vespa olivacea DeGeer, 1773, Mem. Hist. Insect. 3: 582, pl. 29 fig. 9.

Vespa hebraea Fabricius, 1787, Mant. Insect. 1: 292.

Vespa undata Olivier, 1792, Encyc. Méthod. 6: 684.

Vespa macaensis Fabricius, 1793, Entomol. Syst. 2: 259.

? Polistes flavipennis de Saussure, 1853, Ét. Fam. Vesp. 2: 52.

Polistes hebreus [!]; Radoszkowski, 1871, Horae Entomol. Soc. Ross. 8: 195.

Polistes olivaceous [!]; Liu, 1936-37, Peking Nat. Hist. Bull. 11 (3): 211. - Sonan, 1943, Trans. Nat. Hist. Soc. Taiwan 33 (242-243): 467, 477.

Distribution: Madagascar; Réunion; Mauritius; Tanzania: Zanzibar; Amirantes; Seychelles; Chagos Archipelago; Egypt; Oman; Iran; Afghanistan; India: Jammu and Kashmir, Assam, Himachal Pradesh, Karnataka, Madhya Pradesh, Manipur, Meghalaya, Sikkim, Tripura, Uttaranchal, Uttar Pradesh, West Bengal; Sri Lanka; Nepal; Myanmar; southern China; (?) Japan: Okinawa; Philippines; Vietnam; Laos; Cambodia; Thailand; Malaysia; Singapore; Indonesia: Sumatra, Borneo; Marianas; New Caledonia; Fiji; Tongas; Samoa; Society Is.: Tahiti; Tuamotu Archipelago: Fakarava; Marquesas: Tahuata, Hiva Oa; introduced Chile: Easter Is.; U.S.A.: Hawaii; Australia: Queensland; New Zealand (Bequaert, 1918, Bull. Am. Mus. Nat. Hist. 39; Dover, 1929, Bull. Raffles Mus. 2; Giordani Soika, 1962, Vidensk. Medd. Dansk Naturh. Foren. 124; Richards, 1978, Soc. Wasps Am.; Richards, 1978, Aust. J. Zool., Suppl. Ser., 61.; Richards, 1985, Fauna Saudi Arabia 6; Das and Gupta, 1989, Orient. Insect. Monogr. 11).

 

penthicus ater Petersen

Polistes penthicus ater Petersen, 1990, Entomol. Scand. 21: 66.

Distribution: Admiralty Is.: Manus.

 

penthicus malaitensis van der Vecht

Polistes penthicus malaitensis van der Vecht, 1972 (1971), Entomol. Essays to commemorate the retirement of Prof. K. Yasumatsu: 92.

Distribution: Solomon Is.: Malaita, Tulagi.

 

penthicus penthicus van der Vecht

Polistes penthicus penthicus van der Vecht, 1972 (1971), Entomol. Essays to commemorate the retirement of Prof. K. Yasumatsu: 91.

Distribution: Solomon Is.: Santa Isabel, Nggela, Tulagi, Guadalcanal.

 

perflavus van der Vecht

Polistes perflavus van der Vecht, 1972 (1971), Entomol. Essays to commemorate the retirement of Prof. K. Yasumatsu: 93.

Distribution: Solomon Is.: Guadalcanal.

 

rothneyi carletoni van der Vecht

Polistes rothneyi carletoni van der Vecht, 1968, Bijdr. Dierk. 38: 100.

Distribution: Pakistan; India: Jammu and Kashmir, Himachal Pradesh, Uttaranchal, West Bengal; Nepal (Das and Gupta, 1989, Orient. Insect. Monogr. 11; Gusenleitner, 2006, Linz. Biol. Beitr. 38 (1): 679).

 

rothneyi engeli van der Vecht

Polistes rothneyi engeli van der Vecht, 1968, Bijdr. Dierk. 38: 108, female - gSumatra, Padangh (Leiden).

Distribution: Indonesia: Sumatra.

 

rothneyi grahami van der Vecht

Polistes rothneyi grahami van der Vecht, 1968, Bijdr. Dierk. 38: 104.

Distribution: China.

 

rothneyi gressitti van der Vecht

Polistes rothneyi gressitti van der Vecht, 1968, Bijdr. Dierk. 38: 106, female - gFormosa, Taihanrokuh (Leiden).

Distribution: Taiwan.

 

rothneyi hainanensis van der Vecht

Polistes rothneyi hainanensis van der Vecht, 1968, Bijdr. Dierk. 38: 106.

Distribution: China: Hainan Is.

 

rothneyi helvenacus van der Vecht

Polistes rothneyi helvenacus van der Vecht, 1968, Bijdr. Dierk. 38: 109.

Distribution: Indonesia: Java.

 

rothneyi ingrami van der Vecht

Polistes rothneyi ingrami van der Vecht, 1968, Bijdr. Dierk. 38: 106.

Distribution: Japan: Okinawa.

 

rothneyi iwatai van der Vecht

Polistes rothneyi iwatai van der Vecht, 1968, Bijdr. Dierk. 38: 104.

Distribution: China; Korea; Japan (Lee, 1982, Econ. Insect. Fauna China Fasc. 30).

 

rothneyi koreanus van der Vecht

Polistes rothneyi koreanus van der Vecht, 1968, Bijdr. Dierk. 38: 104.

Distribution: China; Korea.

 

rothneyi krombeini van der Vecht

Polistes rothneyi krombeini van der Vecht, 1968, Bijdr. Dierk. 38: 100.

Distribution: India: Uttaranchal, Meghalaya, Karnataka, Kerala, Tamil Nadu (Das and Gupta, 1989, Orient. Insect. Monogr. 11; Gusenleitner, 2006, Linz. Biol. Beitr. 38 (1): 679).

 

rothneyi quatei van der Vecht

Polistes rothneyi quatei van der Vecht, 1968, Bijdr. Dierk. 38: 106.

Distribution: Vietnam.

 

rothneyi robinsoni van der Vecht

Polistes rothneyi robinsoni van der Vecht, 1968, Bijdr. Dierk. 38: 106.

Distribution: Malaysia.

 

rothneyi rothneyi Cameron

Polistes rothneyi Cameron, 1900, Ann. Mag. Nat. Hist. (7) 6: 410.

Polistes rufolineatus Cameron, 1900, Ann. Mag. Nat. Hist. (7) 6: 411.

Distribution: India: Delhi, Uttar Pradesh, West Bengal, Meghalaya; Nepal (Das and Gupta, 1989, Orient. Insect. Monogr. 11; Starr, 1992, Bull. Natl. Mus. Nat. Sci. 3; Gusenleitner, 2006, Linz. Biol. Beitr. 38 (1): 679).

 

rothneyi sikkimensis van der Vecht

Polistes rothneyi sikkimensis van der Vecht, 1968, Bijdr. Dierk. 38: 102.

Distribution: India: Sikkim, Bihar, West Bengal, Assam, Meghalaya; Nepal (Das and Gupta, 1989, Orient. Insect. Monogr. 11).

 

rothneyi tibetanus van der Vecht

Polistes rothneyi tibetanus van der Vecht, 1968, Bijdr. Dierk. 38: 102.

Distribution: China: Tibet.

 

rothneyi vechti Das and Gupta

Polistes rothneyi vechti Das and Gupta, 1984, Orient. Insect. 17: 410, nomen nudum.

Polistes rothneyi vechti Das and Gupta, 1989, Orient. Insect. Monogr. 11: 58.

Distribution: India: Manipur; Myanmar.

 

rothneyi yayeyamae Matsumura

Polistes yayeyamae Matsumura, 1908, in Kuroiwa, List Hym. Loochoo: 6, nomen nudum.

Polistes yayeyamae Matsumura, 1911, Thousand Insect. Japan, Suppl. 3: 106, pl. 39 fig. 5.

Polistes yayeyamensis [!]; Yasumatsu, 1935, Annot. Zool. Japan 15: 37.

Polistes jokohamae var. yaeyamae [!]; Starr, 1992, Bull. Natl. Mus. Nat. Sci. 3: 122.

Distribution: Japan: Yaeyama Is. of Ryukyu Is.

 

schach (Fabricius)

Vespa schach Fabricius, 1781, Spec. Insect. 1: 461.

Vespa schuch [!]; Olivier, 1792, Encyc. Méthod. 6: 682.

Distribution: Australia: Northern Territory, Queensland, New South Wales (Richards, 1978, Aust. J. Zool., Suppl. Ser., 61).

 

semiflavus Holmgren

Polistes semiflavus Holmgren, 1868, K. Svenska Fregat. Eugenies Resa Omkring Jorden, Zool. 1, Insect., Hym.: 439.

Distribution: Marianas, Guam (Bequaert and Yasumatsu, 1939, Tenthredo 2 (3)).

 

stenopus brandti van der Vecht

Polistes stenopus brandti van der Vecht, 1972 (1971), Entomol. Essays to commemorate the retirement of Prof. K. Yasumatsu: 98.

Distribution: Solomon Is.: Buka, Bougainville, New Georgia Is., Santa Isabel.

 

stenopus ganonganus van der Vecht

Polistes stenopus ganonganus van der Vecht, 1972 (1971), Entomol. Essays to commemorate the retirement of Prof. K. Yasumatsu: 99.

Distribution: Solomon Is.: Ranongga, Vella Lavella.

 

stenopus manni van der Vecht

Polistes stenopus manni van der Vecht, 1972 (1971), Entomol. Essays to commemorate the retirement of Prof. K. Yasumatsu: 98.

Distribution: Solomon Is.: Malaita.

 

stenopus stenopus van der Vecht

Polistes stenopus stenopus van der Vecht, 1972 (1971), Entomol. Essays to commemorate the retirement of Prof. K. Yasumatsu: 93, 96.

Distribution: Solomon Is.: Nggela, Tulagi, Guadalcanal, Russell Is.

 

tenebricosus Lepeletier

Polistes tenebricosa Lepeletier, 1836, Hist. Nat. Insect. Hym. 1: 529.

Polistes sulcatus Smith, 1852, Trans. Entomol. Soc. London (2) 2: 38, male, female - China: Near Ning-po-fo (London).

Polistes hoplitus de Saussure, 1853, Ét. Fam. Vesp. 2: 55, female - "Indes Orientales" (Genève).

Polistes hoplites de Saussure, 1854, Ét. Fam. Vesp. 2, Errata: 255 (unjustified emendation).

Polistes rugifrons Cameron, 1900, Ann. Mag. Nat. Hist. (7) 6: 412 - India: Meghalaya: Khasi Hills.

Polistes varicornis Cameron, 1905, Tijdschr. Entomol. 48: 67, male (Amsterdam).

Polistes javanicus Cameron, 1905, Tijdschr. Entomol. 48: 68, female - gTjandi near Semarang, Javah (lectotype Amsterdam).

Polistes tenebricosus var. leopoldi Bequaert, 1934, Bull. Mus. R. Hist. Nat. Belg. 10 (28): 9.

Polistes tenebricosus var. (or subsp.) nigrosericans Bequaert, 1940, Trans. Am. Entomol. Soc. 66: 266.

Polistes tenebricosus var. (or subsp.) sibuyanensis Bequaert, 1940, Trans. Am. Entomol. Soc. 66: 266.

Polistes tenebricosus nigrosercans [!]; Yoshikawa, 1962, J. Biol. Osaka City Univ. 13: 38.

Distribution: India: Jammu and Kashmir, Uttaranchal, Sikkim, West Bengal, Assam, Meghalaya; Nepal; China; Taiwan; Vietnam; Myanmar; Indonesia: Sumatra, Java, Bali, Lombok, Flores, Sumba, Borneo; Sulawesi; Philippines: Luzon, Negros, Sibuyan (Bequaert, 1934, Bull. Mus. R. Hist. Nat. Belg. 10 (28). - Das and Gupta, 1984, Orient. Insect. 17. - Starr, 1992, Bull. Natl. Mus. Nat. Sci. 3. - Saito et al., 2005, J. Hym. Res. 14. - Gusenleitner, 2006, Linz. Biol. Beitr. 38 (1): 679. - Kojima et al., 2006, Serangga 11 (1-2): 12).

 

tenuispunctia Kim

Polistes tenuispunctia Kim, 2001, Kor. J. Entomol. 31 (1): 59, female, male - [Korea:] gAenmubong, Gyeonggi-doh (holotype female Seoul); also from numerous other localities.

Distribution: Korea.

 

tepidus batjanensis Cameron

Polistes tepidus race batjanensis Cameron, 1911, Nova Guinea, Zool. 9 (2), Zool.: 186, female - gBatjanh (Amsterdam).

Distribution: Indonesia: Moluccas: Bacan.

 

tepidus malayanus Cameron

Polistes malayanus Cameron, 1906, Nova Guinea, Zool. 5: 60, female, male - gManokwarih (lectotype male Amsterdam).

Polistes tepidus var. minor von Schulthess, 1932, Mém. R. Mus. Hist. Nat. Belg. hors série 4 (5): 39, non Palisot de Beauvois, 1818.

? Polistes greligiosus Cheesm.h Richards, 1973, Rev. Bras. Entomol. 17 (13): 100, nomen nudum.

Distribution: Indonesia: Aru, Kai Is.; New Guinea; Bismarck Archipelago: New Ireland, Djaul, Lavongai; Solomon Is.: Buka to Rennell; Vanuatu; introduced U.S.A.: Hawaii (van der Vecht, 1972 (1971), Entomol. Essays to commemorate the retirement of Prof. K. Yasumatsu; Richards, 1978, Aust. J. Zool., Suppl. Ser., 61; Petersen, 1990, Entomol. Scand. 21).

 

tepidus manusensis Petersen

Polistes tepidus manusensis Petersen, 1990, Entomol. Scand. 21: 63.

Distribution: Admiralty Is.: Manus, Los Negros.

 

tepidus picteti de Saussure

Polistes picteti de Saussure, 1853, Ét. Fam. Vesp. 2: 69.

Distribution: Indonesia: Moluccas: Ambon, Seram; New Guinea (Richards, 1978, Aust. J. Zool., Suppl. Ser., 61).

 

tepidus speciosus du Buysson

Polistes tepidus var. speciosus du Buysson, 1913 (1911), Abh. Senckenberg. Naturforsch. Ges. 34: 227.

Distribution: Indonesia: Moluccas: Ternate, Bacan (Richards, 1978, Aust. J. Zool., Suppl. Ser., 61).

 

tepidus tepidus (Fabricius)

Vespa tepida Fabricius, 1775, Syst. Entomol.: 366.

Distribution: Australia: Queensland, New South Wales, Victoria (Richards, 1978, Aust. J. Zool., Suppl. Ser., 61; Petersen, 1990, Entomol. Scand. 21).

 

tepidus theodori Petersen

Polistes tepidus theodori Petersen, 1990, Entomol. Scand. 21: 64.

Distribution: Indonesia: Kai Is.

 

wattii Cameron

Polistes wattii Cameron, 1900, Ann. Mag. Nat. Hist. (7) 6: 416.

Polistes watti [!]; Dalla Torre, 1904, Genera Insect. 19: 71.

Distribution: Afghanistan; Iran; Iraq; Saudi Arabia; United Arab Emirates; Oman; Mauritius; Pakistan; India: Himachal Pradesh, Punjab, Delhi, Uttaranchal, West Bengal, Meghalaya, Gujarat, Orissa; China (Gusenleitner, 1972 (1971-72), Acta Mus. Morav. 56/57; Richards, 1985, Fauna Saudi Arabia 6; Das and Gupta, 1989, Orient. Insect. Monogr. 11).

 

 

SUBGENUS POLISTELLA ASHMEAD

(phyletic sequence incertae sedis)

 

Polistella Ashmead, 1904, Proc. U. S. Natl. Mus. 28: 133, genus.

Type species: Polistes manillensis de Saussure, 1853, by original designation.  [Misidentification of Protopolybia sedula (de Saussure) = P. exigua (de Saussure).]

Stenopolistes van der van der Vecht, 1972 (1971), Entomol. Essays to commemorate the retirement of Prof. K. Yasumatsu: 101, subgenus of Polistes Latreille.

Type species: Polistes lateritius Smith, 1857, by original designation.

 

adustus Bingham

Polistes adustus Bingham, 1897, Fauna Br. India Hym. 1: 397.

Polistes sordidus Dover, 1925 (1924), J. Proc. Asiat. Soc. Bengal (N. S) 20: 303, nomen nudum.

Polistes adstus [!]; Yoshikawa, 1962, J. Biol. Osaka City Univ. 13: 20.

Distribution: India: Himachal Pradesh, Delhi, Uttaranchal, Sikkim, Meghalaya, West Bengal; Nepal; China (Lee, 1982, Econ. Insect. Fauna China Fasc. 30; Das and Gupta, 1989, Orient. Insect. Monogr. 11).

 

affinis Gusenleitner

Polistes affinis Gusenleitner, 2006, Linz. Biol. Beitr. 38 (1): 671, 672, female (in subgenus Polistella) - gLaos, Phongsaly env., Prov. Phongsaly, 1500 m, 2141-2'N 10206-8'Eh (Oberösterreichischen Landesmuseums Linz).

Distribution: Laos.

 

albocalcaratus du Buysson

Polistes albocalcaratus du Buysson, 1905, Bull. Soc. Entomol. France 1905: 257.

Distribution: Madagascar.

 

angusticlypeus Gusenleitner

Polistes angusticlypeus Gusenleitner, 2006, Linz. Biol. Beitr. 38 (1): 677, 680, female, male (in subgenus Polistella) - gIndia: Shilloug 33km N, Megalaya, 2605'N 9223'Eh [Shillong in Meghalaya] (holotype male Oberösterreichischen Landesmuseums Linz); also from elsewhere in Meghalaya.

Distribution: India: Meghalaya.

 

aquilinus du Buysson

Polistes aquilinus du Buysson, 1905, Bull. Soc. Entomol. France 1905: 256.

Distribution: Sierra Leone; Liberia; Cameroon; Congo Republic; Congo (Bequaert, 1938, Rev. Zool. Bot. Afr. 31: 134).

 

assamensis Bingham

Polistes assamensis Bingham, 1897, Fauna Br. India Hym. 1: 397.

Distribution: India: Sikkim, Assam (Das and Gupta, 1984, Orient. Insect. 17).

 

asterope arvidi Petersen

Polistes asterope arvidi Petersen, 1990, Entomol. Scand. 21: 61.

Distribution: Bismarck Archipelago: Lavongai.

 

asterope asterope Cameron

Polistes asterope Cameron, 1901, Proc. Zool. Soc. 1901: 243.

Distribution: Bismarck Archipelago: New Britain (Petersen, 1990, Entomol. Scand. 21).

 

asterope narupi Petersen

Polistes asterope narupi Petersen, 1990, Entomol. Scand. 21: 61.

Distribution: Bismarck Archipelago: New Ireland.

 

bambusae bambusae Richards

Polistes bambusae Richards, 1978, Aust. J. Zool., Suppl. Ser., 61: 118.

Distribution: New Guinea.

 

bambusae humboldti Richards

Polistes bambusae humboldti Richards, 1978, Aust. J. Zool., Suppl. Ser., 61: 120.

Distribution: New Guinea.

 

bequaerti von Schulthess

Polistes bequaerti von Schulthess, 1921, Entomol. Mitt. 10: 123.

Distribution: Congo; Zambia (Bequaert, 1938, Rev. Zool. Bot. Afr. 31: 139).

 

bituberculatus du Buysson

Polistes bituberculatus du Buysson, 1905, Bull. Soc. Entomol. France 1905: 257.

Distribution: Madagascar.

 

buruensis Petersen

Polistes buruensis Petersen, 1990, Entomol. Scand. 21: 58.

Distribution: Indonesia: Moluccas: Buru.

 

callimorpha de Saussure

Polistes callimorpha de Saussure, 1853, Ét. Fam. Vesp. 2: 71.

Distribution: Indonesia: Timor, Sumba, (?) Kai Is. (Saito et al., 2005, J. Hym. Res. 14).

 

capnodes capnodes van der Vecht

Polistes capnodes van der Vecht, 1972 (1971), Entomol. Essays to commemorate the retirement of Prof. K. Yasumatsu: 102.

Distribution: Solomon Is.: Guadalcanal.

 

capnodes incomptus van der Vecht

Polistes capnodes incomptus van der Vecht, 1972 (1971), Entomol. Essays to commemorate the retirement of Prof. K. Yasumatsu: 104.

Distribution: Solomon Is.: Malaita.

 

capnodes quadrifasciatus van der Vecht

Polistes capnodes quadrifasciatus van der Vecht, 1972 (1971), Entomol. Essays to commemorate the retirement of Prof. K. Yasumatsu: 104.

Distribution: Solomon Is.: Kolombangara, New Georgia.

 

clavicornis van der Vecht

Polistes clavicornis van der Vecht, 1972 (1971), Entomol. Essays to commemorate the retirement of Prof. K. Yasumatsu: 105.

Distribution: Solomon Is.: Bougainville, Choiseul, Santa Isabel.

 

contrarius Cheesman

Polistes contrarius Cheesman, 1951, Ann. Mag. Nat. Hist. (12) 4: 985.

Distribution: New Guinea.

 

dawnae Dover and Rao

Polistes dawnae Dover and Rao, 1922, J. Proc. Asiat. Soc. Bengal (n. s.) 18: 248.

Distribution: Myanmar.

 

defectivus Gerstaecker

Polistes defectivus Gerstaecker, 1871, Arch. Naturgesch. 37: 351.

Polistes waldoi Dover, 1926, Entomologist 59: 36.

Distribution: Congo; Ethiopia; Tanzania; Malawi; Zimbabwe; Mozambique; South Africa: Eastern Transvaal, Kwazulu-Natal, Western Cape Prov. (Bequaert, 1938, Rev. Zool. Bot. Afr. 31: 148; Giordani Soika, 1952, Riv. Biol. Colon. 11).

 

delhiensis Das and Gupta

Polistes delhiensis Das and Gupta, 1984, Orient. Insect. 17: 410, nomen nudum.

Polistes delhiensis Das and Gupta, 1989, Orient. Insect. Monogr. 11: 63.

Distribution: India: Delhi.

 

diakonovi Kostylev

Polistes diakonovi Kostylev, 1940, Bull. Soc. Nat. Moscou, Sect. Biol 49: 139.

Distribution: Russia: Ussuri.

 

ebsohinus Sonan

Polistes mandarinus var. ebsohinus Sonan, 1943, Trans. Nat. Hist. Soc. Taiwan 33 (242-243): 483.

Distribution: Taiwan (Starr, 1992, Bull. Natl. Mus. Nat. Sci. 3).

 

elegans Smith

Polistes elegans Smith, 1859, J. Proc. Linn. Soc., Zool. 3 (11): 169.

Distribution: Indonesia: Aru, Kai Is.; New Guinea (Cheesman, 1951, Ann. Mag. Nat. Hist. (12) 4).

 

ellenbergi du Buysson

Polistes ellenbergi du Buysson, 1908, Bull. Soc. Entomol. France 1908: 64.

Distribution: Liberia; Cameroon; Equatorial Guinea; Congo Republic (Bequaert, 1918, Bull. Am. Mus. Nat. Hist. 39; Bequaert, 1938, Rev. Zool. Bot. Afr. 31: 149).

 

ephippium Cameron

Polistes ephippium Cameron, 1900, Ann. Mag. Nat. Hist. (7) 6: 415.

Distribution: India: Meghalaya (Das and Gupta, 1989, Orient. Insect. Monogr. 11).

 

extraneus Kirby

Polistes extraneus Kirby, 1883, Proc. Zool. Soc. London 1883: 344.

Distribution: Indonesia: Maru.

 

fastidiosus de Saussure

Polistes fastidiosus de Saussure, 1853, Ét. Fam. Vesp. 2: 60.

Distribution: Senegal; Guinea-Bissau; Ivory Coast; Benin; Burkina Faso; Nigeria; Congo; Sudan; Somalia; Saudi Arabia; Yemen; Kenya; Tanzania, including Zanzibar; Zambia; Mozambique; South Africa: Transvaal, Cape Prov. (Bequaert, 1938, Rev. Zool. Bot. Afr. 31: 140; Richards, 1985, Fauna Saudi Arabia 6).

 

formosanus Sonan

Polistes formosanus Sonan, 1927, Trans. Nat. Hist. Soc. Formosa 17: 122.

Polistes japonicus var. formosana form koshunensis Sonan, 1938, Arbeit. Morphol. Taxon. Entomol. Berlin-Dahlem 5: 68.  Unavailable under Art. 45.5 of the International Code of Zoological Nomenclature.

Polistes japonicus var. fomosanus [!]; Kuo and Yeh, 1987, J. Natl. Chiayi Inst. Agric. 16: 83.

Distribution: Japan, Ryukyu Is.; Taiwan.

 

flavobilineatus (Cameron)

Icaria flavo-bilineata Cameron, 1902, J. Straits Asiat. Soc. 37: 102.

Distribution: Myanmar; Thailand; Malaysia, Sarawak (Dover, 1931, J. Fed. Malay Mus. 16).

 

haugi du Buysson

Polistes haugi du Buysson, 1906, Bull. Soc. Entomol. France 1906: 189.

Distribution: Liberia; Nigeria; Cameroon; Equatorial Guinea; Congo Republic; Congo (Bequaert, 1938, Rev. Zool. Bot. Afr. 31: 149).

 

horrendus Gusenleitner

Polistes horrendus Gusenleitner, 2006, Linz. Biol. Beitr. 38 (1): 671, female (in subgenus Polistella) - gLaos, Phongsaly env., Prov. Phongsaly, 1500 m, 2141-2'N 10206-8'Eh (Oberösterreichischen Landesmuseums Linz).

Distribution: Laos.

 

huisunensis Kuo

Polistes huisunensis Kuo, 1987, in Kuo and Yeh, J. Natl. Chiayi Inst. Agric. 16: 80.

Distribution: Taiwan.

 

humilis centrocontinentalis Giordani Soika

Polistes humilis centrocontinentalis Giordani Soika, 1975, Boll. Soc. Entomol. Ital. 107: 20.

Distribution: central Australia.

 

humilis humilis (Fabricius)

Vespa humilis Fabricius, 1781, Spec. Insect. 1: 461.

Polistes tasmaniensis de Saussure, 1853, Ét. Fam. Vesp. 2: 66.

Polistes tricolor de Saussure, 1853, Ét. Fam. Vesp. 2: 67, non Fabricius, 1775.

Polistes humilis pseudoschach Giordani Soika, 1975, Boll. Soc. Entomol. Ital. 107: 21.

Distribution: Australia: Queensland, New South Wales, A. C. T., Victoria, South Australia; introduced New Zealand (Richards, 1978, Aust. J. Zool., Suppl. Ser., 61).

 

humilis synoecus de Saussure

Polistes synoecus de Saussure, 1853, Ét. Fam. Vesp. 2: 65.

Polistes variabilis var. reginae Meade-Waldo, 1911, Ann. Nat. Hist. (8) 7: 101.

Polistes humilis xanthorrhoicus Giordani Soika, 1975, Boll. Soc. Entomol. Ital. 107: 21.

Polistes humilis clarior Giordani Soika, 1975, Boll. Soc. Entomol. Ital. 107: 22.

Polistes humilis xanthorhoicus [!]; Richards, 1978, Aust. J. Zool., Suppl. Ser., 61: 46.

Distribution: Australia: Northern Territory, Queensland, introduced Western Australia; Society Is.: Moorea, Tahiti (Richards, 1978, Aust. J. Zool., Suppl. Ser., 61).

 

intermedius Kojima

Polistes medius Kojima, 1988, in Kojima and Kojima, J. Aust. Entomol. Soc. 27: 74, non Palisot de Beauvois, 1818.

Polistes intermedius Kojima, 1996, in Carpenter, Am. Mus. Novitat. 3188: 1, 12.  Replacement name for medius Kojima, 1988, non Palisot de Beauvois, 1818.

Distribution: New Guinea.

 

japonicus de Saussure

Polistes japonicus de Saussure, 1858, Rev. Mag. Zool. (2) 10: 260, female - gle Japon.h

Polistes erythrocerus Cameron, 1900, Ann. Mag. Nat. Hist. (7) 6: 418.

Polistes japoniaus [!]; Kuo and Yeh, 1987, J. Natl. Chiayi Inst. Agric. 16: 81.

Polistes hengchunensis Kuo, 1987, in Kuo and Yeh, J. Natl. Chiayi Inst. Agric. 16: 81.

Polistes hengchun [!]; Kuo and Yeh, 1987, J. Natl. Chiayi Inst. Agric. 16: 82.

Polistes shekouensis Kuo, 1987, in Kuo and Yeh, J. Natl. Chiayi Inst. Agric. 16: 82.

Distribution: China; Korea; Japan; Taiwan (Kim, 1980, Distr. Atlas Insect. Korea, Ser. 3, Hym. Dipt.; Lee, 1982, Econ. Insect. Fauna China Fasc. 30; Starr, 1992, Bull. Natl. Mus. Nat. Sci. 3).

 

khasianus Cameron

Polistes khasianus Cameron, 1900, Ann. Mag. Nat. Hist. (7) 6: 413.

Distribution: India: Haryana, Meghalaya, West Bengal (Das and Gupta, 1989, Orient. Insect. Monogr. 11; Gusenleitner, 2006, Linz. Biol. Beitr. 38 (1): 681).

 

laevigatissimus Giordani Soika

Polistes laevigatissimus Giordani Soika, 1975, Boll. Soc. Entomol. Ital. 107: 25.

Distribution: Australia: Western Australia.

 

lateritius Smith

Polistes lateritius Smith, 1857, Cat. Hym. Br. Mus. 5, Vesp.: 103.

Polistes colonicus Smith, 1860, Proc. Linn. Soc. London, Zool. 5 Suppl.: 129.

Polistes lateritus [!]; Yoshikawa, 1962, J. Biol., Osaka Univ. 13: 30.

Distribution: Indonesia: Moluccas: Ambon, Seram, Saparua (Petersen, 1990, Entomol. Scand. 21).

 

latinis Das and Gupta

Polistes latinis Das and Gupta, 1984, Orient. Insect. 17: 412, nomen nudum.

Polistes latinis Das and Gupta, 1989, Orient. Insect. Monogr. 11: 79.

Distribution: India: Andaman Is.

 

lepcha Cameron

Polistes khasianus Cameron, 1900, Ann. Mag. Nat. Hist. (7) 6: 415, non Cameron, 1900: 413.

Polistes lepcha Cameron, 1900, Ann. Mag. Nat. Hist. (7) 6: 506.  Replacement name for khasianus Cameron, 1900: 415, non Cameron, 1900: 413.

Polistes klepcha [!]; Yoshikawa, 1962, J. Biol. Osaka City Univ. 13: 30.

Distribution: India: Haryana, Meghalaya, Manipur (Das and Gupta, 1989, Orient. Insect. Monogr. 11).

 

lycus Cameron

Polistes lycus Cameron, 1901, Proc. Zool. Soc. 1901: 242.

Distribution: Bismarck Archipelago: New Britain, New Ireland (Petersen, 1990, Entomol. Scand. 21).

 

madecassus de Saussure

Polistes madecassus de Saussure, 1853, Ét. Fam. Vesp. 2: 51.

Distribution: Madagascar.

 

madiburensis von Schulthess

Polistes madiburensis von Schulthess, 1921, Entomol. Mitt. 10: 124.

Polistes maniburesis [!]; Yoshikawa, 1962, J. Biol., Osaka Univ. 13: 31.

Distribution: Tanzania; Zambia; Zimbabwe (Bequaert, 1938, Rev. Zool. Bot. Afr. 31: 149).

 

mandarinus de Saussure

Polistes mandarinus de Saussure, 1853, Ét. Fam. Vesp. 2: 58.

Polistes megei Pérez, 1905, Bull. Mus. Hist. Nat. Paris 2: 83.

Polistes mandariniensis [!]; Zimmermann, 1931, Zeitschr. Morphol. Ökol. Tiere 22: 230.

Polistes mandarinius [!]; Ma, 1936, Entomol. Phytopath. 4: 65.

Distribution: China; Korea (Liu, 1937, Peking Nat. Hist. Bull. 11 (3); Kim, 1980, Distr. Atlas Insect. Korea, Ser. 3, Hym. Dipt.; Kojima, 1997, Nat. Hist. Bull. Ibaraki Univ. 1).

 

meadeanus (von Schulthess)

Parapolybia meadeana von Schulthess, 1913, Mitt. Schweiz. Entomol. Ges. 12: 161.

Polybia andrei du Buysson, 1913, Bull. Soc. Entomol. France 1913: 296.

Polybia shelfordi du Buysson, 1913, Bull. Soc. Entomol. France 1913: 297.

Distribution: Malaysia; Indonesia: Borneo (van der Vecht, 1966, Zool. Verh., Leiden 82).

 

melanopterus Cameron

Polistes melanopterus Cameron, 1911, Nova Guinea, Zool. 9 (2), Zool.: 187, female - gLorentz Riverh (lectotype Amsterdam).

Distribution: New Guinea.

 

mertoni du Buysson

Polistes mertoni du Buysson, 1913 (1911), Abh. Senckenberg. Naturforsch. Ges. 34: 227.

Distribution: Indonesia: Aru.

 

nigerrimus Gusenleitner

Polistes nigerrimus Gusenleitner, 2006, Linz. Biol. Beitr. 38 (1): 671, 674, female (in subgenus Polistella) - gVietnam, Phongsaly [error], Tam Dao, Vinh phu Provinceh (Oberösterreichischen Landesmuseums Linz).

Distribution: Vietnam.

 

nigrifrons nigrifrons Smith

Polistes nigrifrons Smith, 1859, J. Proc. Linn. Soc., Zool. 3 (11): 168.

Distribution: Indonesia: Aru, Waigeo; New Guinea.

 

nigrifrons obdurus Cheesman

Polistes nigrifrons obdurus Cheesman, 1951, Ann. Mag. Nat. Hist. (12) 4: 989.

Distribution: New Guinea.

 

nigrifrons stotherti Meade-Waldo

Polistes stotherti Meade-Waldo, 1916 (1915), Br. Ornithol. Union Exped. Dutch New Guinea 1: 14.

Distribution: New Guinea.

 

nigritarsis Cameron

Polistes nigritarsis Cameron, 1900, Ann. Mag. Nat. Hist. (7) 6: 413.

Distribution: India: West Bengal, Meghalaya, Kerala, Karnataka, Tamil Nadu (Das and Gupta, 1989, Orient. Insect. Monogr. 11; Gusenleitner, 2006, Linz. Biol. Beitr. 38 (1): 683).

 

nipponensis Pérez

Polistes nipponensis Pérez, 1905, Bull. Mus. Hist. Nat. Paris 2: 82.

Polistes yamanakai Sonan, 1937, Trans. Nat. Hist. Soc. Formosa 27 (166): 169.

Distribution: China; Japan (Kojima, 1997, Nat. Hist. Bull. Ibaraki Univ. 1).

 

occultus Kojima

Polistes occultus Kojima, 1988, in Kojima and Kojima, J. Aust. Entomol. Soc. 27: 76.

Distribution: New Guinea.

 

opacus Gusenleitner

Polistes opacus Gusenleitner, 2006, Linz. Biol. Beitr. 38 (1): 677, 681, female, male (in subgenus Polistella) - gIndia: Paykong, W-Sikkim, ... 2715'N 8835'Eh (holotype female Stuttgart); also from West Bengal.

Distribution: India: Sikkim, West Bengal.

 

philippinensis funebris Bequaert

Polistes sagittarius var. (or subsp.) funebris Bequaert, 1940, Trans. Am. Entomol. Soc. 66: 269.

Distribution: Philippines: Luzon.

 

philippinensis philippinensis de Saussure

Polistes philippinensis de Saussure, 1853, Ét. Fam. Vesp. 2: 58.

Polistes phillipinensis [!]; Meade-Waldo, 1911, Ann. Mag. Nat. Hist. (8) 7: 102.

Distribution: Philippines: Mindanao (Bequaert, 1940, Trans. Am. Entomol. Soc. 66: 265-273).

 

praenotatus Kohl

Polistes praenotatus Kohl, 1908, Denkschr. Akad. Wiss., Wien 81: 311.

Distribution: Solomon Is.

 

quadricingulatus Gusenleitner

Polistes quadricingulatus Gusenleitner, 2006, Linz. Biol. Beitr. 38 (1): 677, 683, female (in subgenus Polistella) - gN-India: Bhimtal, 1500mh (Linz); also from Kashmir, Uttar Pradesh; and Nepal.

Distribution: India: Jammu and Kashmir, Uttaranchal, Uttar Pradesh; Nepal.

 

riekii Richards

Polistes riekii Richards, 1978, Aust. J. Zool., Suppl. Ser., 61: 23.

Distribution: Australia: Queensland.

 

rubellus Gusenleitner

Polistes rubellus Gusenleitner, 2006, Linz. Biol. Beitr. 38 (1): 677, 684, female (in subgenus Polistella) - gIndia: Yuksam 8 km W-Sikkimh (Stuttgart).

Distribution: India: Sikkim; Nepal.

 

rufodorsalis Yamane and Kusigemati

Polistes rufodorsalis Yamane and Kusigemati, 1985, Occas. Pap. Kagoshima Univ. Res. Center S. Pacif. 5: 77.

Distribution: Solomon Is.: Guadalcanal.

 

sagittarius de Saussure

Polistes sagittarius de Saussure, 1853, Ét. Fam. Vesp. 2: 56.

Polistes sagittarius var (or subsp.) indonesicus Bequaert, 1940, Trans. Am. Entomol. Soc. 66: 267.

Polistes sagittarium [!]; Das and Gupta, 1989, Orient. Insect. Monogr. 11: 67.

Polistes strigosus baliensis Gusenleitner, 2000, Linzer Biol. Beitr. 37 (2): 939, male, female (in subgenus Polistella) - gIndonesia, Bali, Nusa Dua Beachh (holotype male Oberösterreichischen Landesmuseum, Linz); also from Celik.

Distribution: India: Himachal Pradesh, Delhi, Kerala, Uttaranchal, Sikkim, West Bengal, Assam, Nagaland, Manipur, Tripura; Nepal; China; Hong Kong; Myanmar; Thailand; Malaysia; Singapore; Indonesia: Sumatra, Borneo, Java, Bali, Lombok, Sulawesi, Flores, Sumbawa, Sumba, Sulawesi; Philippines: Palawan (Das and Gupta, 1989, Orient. Insect. Monogr. 11. - Kojima and Yamane, 1990, in Sakagami et al., Nat. Hist. Soc. Wasps Bees in Eq. Sumatra. - Saito et al., 2005, J. Hym. Res. 14. - Kojima et al., 2006, Serangga 11 (1-2): 11).

 

santoshae Das and Gupta

Polistes virendrai Das and Gupta, 1984, Orient. Insect. 17: 415, nomen nudum.

Polistes santoshae Das and Gupta, 1989, Orient. Insect. Monogr. 11: 72.

Distribution: India: Sikkim, Meghalaya; Nepal.

 

saussurei Dalla Torre

Polistes incertus de Saussure, 1890, in Grandidier, Hist. Madagascar 20, Hym.: 145, non Cresson, 1865.

Polistes saussurei Dalla Torre, 1894, Cat. Hym. 9: 134.  Replacement name for incertus de Saussure, 1890, non Cresson, 1865.

Distribution: Madagascar; Comoros (Bequaert, 1918, Bull. Am. Mus. Nat. Hist. 39).

 

sgarambus Giordani Soika

Polistes sgarambus Giordani Soika, 1975, Boll. Soc. Entomol. Ital. 107: 24.

Distribution: Australia: Northern Territory (Richards, 1978, Aust. J. Zool., Suppl. Ser., 61).

 

shirakii Sonan

Polistes shirakii Sonan, 1943, Trans. Nat. Hist. Soc. Taiwan 33 (242-243): 481.

Polistes shiraki [!]; Yoshikawa, 1962, J. Biol., Osaka Univ. 13: 36.

Distribution: Taiwan.

 

sikorae de Saussure

Polistes sikorae de Saussure, 1900, Abh. Senckenberg. Naturforsch. Ges. 26: 233.

Polistes sikorae var. obscura de Saussure, 1900, Abh. Senckenberg. Naturforsch. Ges. 26: 234, non de Saussure, 1863.

Distribution: Madagascar.

 

similis Das and Gupta

Polistes similis Das and Gupta, 1984, Orient. Insect. 17: 413, nomen nudum.

Polistes similis Das and Gupta, 1989, Orient. Insect. Monogr. 11: 72.

Distribution: India: Meghalaya.

 

simulatus halmaheirensis Petersen

Polistes simulatus halmaheirensis Petersen, 1990, Entomol. Scand. 21: 58.

Distribution: Indonesia: Moluccas: Halmahera.

 

simulatus morotaiensis Petersen

Polistes simulatus morotaiensis Petersen, 1990, Entomol. Scand. 21: 58.

Distribution: Indonesia: Moluccas: Morotai.

 

simulatus obiensis Petersen

Polistes simulatus obiensis Petersen, 1990, Entomol. Scand. 21: 57.

Distribution: Indonesia: Moluccas: Obi.

 

simulatus simulatus Smith

Polistes simulatus Smith, 1860, J. Proc. Linn. Soc., Zool. 5 Suppl.: 130.

Distribution: Indonesia: Moluccas: Bacan, Kayoa (Petersen, 1990, Entomol. Scand. 21).

 

smithii neavei von Schulthess

Polistes smithii var. neavei von Schulthess, 1921, Entomol. Mitt. 10: 126.

Distribution: Sierra Leone; Congo; Tanzania, including Zanzibar; Mozambique; Zambia (Bequaert, 1938, Rev. Zool. Bot. Afr. 31: 149).

 

smithii smithii de Saussure

Polistes smithii de Saussure, 1853, Ét. Fam. Vesp. 2: 60.

Distribution: Senegal; Congo; Tanzania, including Zanzibar; Zimbabwe; South Africa: Kwazulu-Natal (Bequaert, 1938, Rev. Zool. Bot. Afr. 31: 146; Richards and Richards, 1951, Trans. R. Entomol. Soc. London 102).

 

snelleni de Saussure

Polistes snelleni de Saussure, 1862, Stettin. Entomol. Ztg. 23: 140, female - gJaponh (lectotype Leiden).

Polistes puncticollis Morawitz, 1892, Horae Entomol. Soc. Ross. 26: 150.

Polistes snelleni form nigrihumerus Uchida, 1936, Biogeographica, Tokyo 1 (2): 69.

Distribution: Russia: Siberia; China; Korea; Japan (Kim, 1980, Distr. Atlas Insect. Korea, Ser. 3, Hym. Dipt.).

 

stigma alagari Petersen

Polistes stigma alagari Petersen, 1987, Entomol. Scand. 18: 246.

Distribution: Philippines: Tawitawi, Palawan.

 

stigma bernardii Le Guillou

Polistes bernardii Le Guillou, 1841, Rev. Mag. Zool. 5: 325.

Polistes bernardii richardsi Giordani Soika, 1975, Boll. Soc. Entomol. Ital. 107: 23.

Polistes bernardii duplicinctus Richards, 1978., Aust. J. Zool., Suppl. Ser., 61: 28.

Polistes bernardii insulae Richards, 1978, Aust. J. Zool., Suppl. Ser., 61: 30.

Distribution: New Guinea; Australia: Western Australia, Northern Territory, Queensland; Society Is.: Tahiti (Cheesman, 1928, Ann. Mag. Nat. Hist. (10) 1; Richards, 1978, Aust. J. Zool., Suppl. Ser., 61; Petersen, 1987, Entomol. Scand. 18).

 

stigma dubius de Saussure

Polistes dubius de Saussure, 1867, Reise Novara, Zool. 2 (1), Hym.: 20, pl. I fig. 15, female, male - gManilah (Wien).

Distribution: Philippines: Luzon, Calicoan Is. (Petersen, 1987, Entomol. Scand. 18).

 

stigma galatheae Petersen

Polistes stigma galatheae Petersen, 1987, Entomol. Scand. 18: 239.

Distribution: India: South Nicobar Is.

 

stigma goestai Petersen

Polistes stigma goestai Petersen, 1987, Entomol. Scand. 18: 255.

Distribution: Bismarck Archipelago: Mussau.

 

stigma jani Petersen

Polistes stigma jani Petersen, 1987, Entomol. Scand. 18: 256.

Distribution: Admiralty Is.: Manus.

 

stigma maculipennis de Saussure

Polistes maculipennis de Saussure, 1853, Ét. Fam. Vesp. 2: 61.

Polistes multipictus Smith, 1860, Proc. Linn. Soc. London, Zool. 5 Suppl.: 130, non Haliday, 1836.

Polistes stigma var. thoracica du Buysson, 1913 (1911), Abh. Senckenberg. Naturforsch. Ges. 34: 228, non Fox, 1898.

Polistes comis Cheesman, 1951, Ann. Mag. Nat. Hist. (12) 4: 991.

Distribution: New Guinea; Indonesia: Aru, Moluccas: Ambon (Petersen, 1987, Entomol. Scand. 18).

 

stigma madsi Petersen

Polistes stigma madsi Petersen, 1987, Entomol. Scand. 18: 254.

Distribution: Bismarck Archipelago: Djaul.

 

stigma manillensis de Saussure

Polistes manillensis de Saussure, 1853, Ét. Fam. Vesp. 2: 70.

Polistes manillesis [!]; Yoshikawa, 1962, J. Biol., Osaka Univ. 13: 31.

Distribution: Philippines: Luzon, Leyte (Kojima, 1984, Kontyû 52; Petersen, 1987, Entomol. Scand. 18).

 

stigma nebulosus Yamane and Kusigemati

Polistes stigma nebulosus Yamane and Kusigemati, 1985, Occas. Pap. Kagoshima Univ. Res. Center S. Pacif. 5: 77.

Distribution: Solomon Is.: Guadalcanal; Fiji.

 

stigma novarae de Saussure

Polistes novarae de Saussure, 1867, Reise Novara, Zool. 2 (1), Hym.: 19, pl. 1 figs. 13-14, male, female - gNicobaricus Insulis, in Sinu Bengalensi = Telluschongh (Wien); also from two other localities; and Australia.

Distribution: India: North Nicobar Is. (Petersen, 1987, Entomol. Scand. 18).

 

stigma papuanus Schulz

Polistes marginalis papuanus Schulz, 1905, Berlin Entomol. Zeitschr. 49: 231.

Distribution: Bismarck Archipelago: New Britain; Duke of York; New Ireland (Petersen, 1987, Entomol. Scand. 18).

 

stigma pouli Petersen

Polistes stigma pouli Petersen, 1987, Entomol. Scand. 18: 252.

Distribution: Bismarck Archipelago: Lavongai.

 

stigma sauiensis Petersen

Polistes stigma sauiensis Petersen, 1987, Entomol. Scand. 18: 240.

Distribution: India: Car Nicobar Is.

 

stigma stigma (Fabricius)

Vespa stigma Fabricius, 1793, Entomol. Syst. 2: 275.

Distribution: Thailand; Malaysia; Singapore; Indonesia: Sumatra, Java, Bali, Lombok; Taiwan (Dover, 1929, Bull. Raffles Mus. 2; Petersen, 1987, Entomol. Scand. 18; Kojima and Yamane, 1990, in Sakagami et al., Nat. Hist. Soc. Wasps Bees in Eq. Sumatra; Starr, 1992, Bull. Natl. Mus. Nat. Sci. 3).

 

stigma svendi Petersen

Polistes stigma svendi Petersen, 1987, Entomol. Scand. 18: 257.

Distribution: Hermit Is.: Luf.

 

stigma tamula (Fabricius)

Vespa tamula Fabricius, 1798, Suppl. Entomol. Syst.: 263.

Distribution: India: Himachal Pradesh, Delhi, Bihar, West Bengal, Maharashtra, Karnataka, Kerala, Tamil Nadu; Sri Lanka (Das and Gupta, 1989, Orient. Insect. Monogr. 11; Gusenleitner, 2006, Linz. Biol. Beitr. 38 (1): 686).

 

stigma townsvillensis Giordani Soika

Polistes townsvillensis Giordani Soika, 1975, Boll. Soc. Entomol. Ital. 107: 22.

Polistes townsvillensis austrinus Richards, 1978, Aust. J. Zool., Suppl. Ser., 61: 37.

Distribution: Australia: Queensland, New South Wales Victoria (Petersen, 1987, Entomol. Scand. 18).

 

stigma tualensis Petersen

Polistes stigma tualensis Petersen, 1987, Entomol. Scand. 18: 243.

Distribution: Indonesia: Kai Is.

 

strigosus atratus Das and Gupta

Polistes strigosus atratus Das and Gupta, 1984, Orient. Insect. 17: 415, nomen nudum.

Polistes strigosus atratus Das and Gupta, 1989, Orient. Insect. Monogr. 11: 81.

Distribution: India: Delhi, Uttar Pradesh, Sikkim, Bihar, West Bengal, Assam, Manipur, Tripura; Nepal.

 

strigosus mimus Bequaert

Polistes strigosus var. (or subsp.) mimus Bequaert, 1940, Trans. Am. Entomol. Soc. 66: 272.

Polistes strigosus minus [!]; Kojima and Yamane, 1984, Rep. Fac. Sci., Kagoshima Univ. (Earth Sci. Biol) 17: 119.

Distribution: Nepal; Malaysia: Sabah; Philippines: Sibuyan, Luzon, Mindanao (Yamane and Yamane, 1979, Insect. Matsum., N. S. 15; Kojima and Yamane, 1984, Rep. Fac. Sci., Kagoshima Univ. (Earth Sci. Biol) 17).

 

strigosus strigosus Bequaert

Polistes strigosus Bequaert, 1940, Trans. Am. Entomol. Soc. 66: 269.

Polistes stigosus [!]; Das and Gupta, 1984, Orient. Insect. 17: 415.

Distribution: China; Taiwan (Sonan, 1943, Trans. Nat. Hist. Soc. Formosa 33: 467-484).

 

takasagonus Sonan

Polistes takasagonus Sonan, 1943, Trans. Nat. Hist. Soc. Taiwan 33 (242-243): 482.

Distribution: Taiwan.

 

tristis Meade-Waldo

Polistes tristis Meade-Waldo, 1911, Ann. Nat. Hist. (8) 7: 101.

Distribution: Kenya.

 

tullgreni Schulz

Polistes inornatus Tullgren, 1904, Arkiv Zool. 1: 456, pl. 25 figs. 15a, b, non Ritsema, 1874.

Polistes tullgreni Schulz, 1906, Spolia Hym.: 229.  Replacement name for inornatus Tullgren, 1904, non Ritsema, 1874.

Distribution: Cameroon; Congo (Bequaert, 1938, Rev. Zool. Bot. Afr. 31: 151).

 

utakwae Meade-Waldo

Polistes utakwae Meade-Waldo, 1916 (1915), Br. Ornithol. Union Exped. Dutch New Guinea 1: 13.

Distribution: New Guinea.

 

variabilis (Fabricius)

Vespa variabilis Fabricius, 1781, Spec. Insect. 1: 466.

Distribution: Australia: Queensland (Richards, 1978, Aust. J. Zool., Suppl. Ser., 61).

 

watutus Kojima

Polistes watutus Kojima, 1988, in Kojima and Kojima, J. Aust. Entomol. Soc. 27: 72.

Distribution: New Guinea.

 

williamsi Petersen

Polistes williamsi Petersen, 1990, Entomol. Scand. 21: 58.

Distribution: Philippines: Palawan, Panay, Negros, Sibuyan, Mindanao, Samar, Luzon, Polillo.

 

xantholeucus van der Vecht

Polistes xantholeucus van der Vecht, 1984, Nat. Hist. Rennell Is., Br. Solomon Is. 8: 171.

Distribution: Solomon Is.: Rennell, San Cristobal.

 

 

SUBGENUS POLISTES LATREILLE

 

Polistes Latreille, 1802, Hist. Nat. Crust. Insect. 3: 363, genus (6 species).

Type species: gPolistes gallica, Fab.h [=Vespa gallica Linnaeus, 1767], by subsequent designation of Latreille, 1810, Consid. Gén. Crust. Arachn. Ins.: 438.

Polystes [!]; Palisot de Beauvois, 1818, Insect. Recueill. Afrique Amérique: pl. 8; du Buysson, 1892, Ann. Soc. Entomol. France 61: 59; H. von Ihering, 1896, Zool. Anz. 19: 452.

Eupolistes Dalla Torre, 1904, Genera Insect. 19: 68, name for gPremiere divisionh of Polistes Latreille in de Saussure, 1853, Ét. Fam. Vesp. 2: 45 (61 species).

Type species: Vespa gallica Linnaeus, 1767, by subsequent designation of Richards, 1973, Rev. Bras. Entomol. 17 (13): 86.

Pseudopolistes Weyrauch, 1937, Zool. Jahrb. (Abt. Syst. Ökol. Geogr. Tiere) 70: 266, 274, genus (3 species). Unavailable; no type species designated.

Sulcopolistes Blüthgen, 1938 (1937), Konowia 16: 273, subgenus of Polistes Latreille.

Type species: Polistes semenowi Morawitz, 1889, by original designation.

Polistula Weyrauch, 1938, Arbeit. Physiol. Angewand. Entomol. 5 (3): 273, genus (5 species). Unavailable; no type species designated.

Polistula Weyrauch, 1939, Arch. Naturgesch. (N. F.) 8 (2): 148, genus.

Type species: Polistes kohli Dalla Torre, 1904 [= Polistes biglumis Linnaeus, 1758], by original designation. Validation by type selection of Polistula Weyrauch, 1938.

Pseudopolistes Weyrauch, 1939, Arch. Naturgesch. (N. F.) 8 (2): 195. Validation by type selection of Pseudopolistes Weyrauch, 1937.

Type species: Polistes sulcifer Zimmermann, 1930, by original designation.

Leptopolistes Blüthgen, 1943, Arch. Naturgesch. (N. F.) 12 (1): 99, 121, subgenus of Polistes Latreille.

Type species: Polistes associus Kohl, 1898, by original designation.

 

africanus Palisot de Beauvois

Polistes africana Palisot de Beauvois, 1818, Insect. Recueill. Afrique Amérique, livr. 12: 207.

Polystes aficana [!]; Palisot de Beauvois, 1818, Insect. Recueill. Afrique Amérique: pl. 8.

Polistes plebejus Gerstaecker, 1871, Arch. Naturgesch. 37: 351.

Polistes inornatus Ritsema, 1874, Tijdschr. Entomol. 17: 177, 203, pl. 11 fig. 8.

Polistes spilophorus Schletterer, 1891, Ann. Soc. Entomol. Belg. 35: 29.

Polistes spiloptera [!]; Kohl, 1894, Ann. Naturh. Hofmus., Wien 9: 343.

Polistes spliophorus [!]; Yoshikawa, 1962, J. Biol., Osaka Univ. 13: 31.

Distribution: Senegal; Liberia; Nigeria; Cameroon; São Tomé; Congo; Kenya; Somalia; Tanzania, including Zanzibar; Comoros; South Africa: Kwazulu-Natal, Western Cape Prov. (Bequaert, 1938, Rev. Zool. Bot. Afr. 31: 142; Guiglia, 1940, Boll. Lab. Zool. Gen. Agrar. Fac. Agrar. Portici 31; Giordani Soika, 1961, South Afr. Anim. Life 8).

 

albicinctus de Saussure

Polistes albicinctus de Saussure, 1890, in Grandidier, Hist. Madagascar 20, Hym.: 148.

Polistes albocinctus [!]; de Saussure, 1900, Abh. Senckenberg. Naturforsch. Ges. 36: 234.

Distribution: Madagascar (Giordani Soika, 1991, Lavori Soc. Ven. Sci. Nat. 16).

 

associus Kohl

Polistes associa Kohl, 1898, Ann. Naturh. Hofmus., Wien 13: 89.

Distribution: central and southern Europe; Turkey; Israel; Azerbaijan; India: Jammu and Kashmir; China (Guiglia, 1972, Guêpes Soc. Europe Occident. Septentrion.; Das and Gupta, 1989, Orient. Insect. Monogr. 11; Gusenleitner, 1995, Linzer Biol. Beitr. 27).

 

atrimandibularis Zimmermann

Polistes atrimandibularis Zimmermann, 1930, Mitt. Zool. Mus. Berlin 15: 611.

Polistes atrimandibularis albidus Blüthgen, 1957, Rev. Fac. Sci. Univ. Istanbul, Ser. B, 22 (3): 164.

Distribution: central and southern Europe; Turkey; Iran; Armenia; Egypt (Guiglia, 1972, Guêpes Soc. Europe Occident. Septentrion.; Yildrim and Özbek, 1993, Türk. Entomol. Derg. 17).

 

badius Gerstaecker

Polistes badia Gerstaecker, 1871, Arch. Naturgesch. 37: 351.

Distribution: Senegal; Guinea-Bissau; Gambia; Togo; Gabon; Congo; Tanzania, including Zanzibar, Wanga; Kenya; Somalia; Ethiopia; Sudan; Yemen (Bequaert, 1938, Rev. Zool. Bot. Afr. 31: 143; Guiglia, 1939, Mem. Soc. Entomol. Ital. 18; Guiglia, 1940, Boll. Lab. Zool. Gen. Agrar. Fac. Agrar. Portici 31; Giordani Soika, 1952, Riv. Biol. Colon. 11; Giordani Soika, 1957, Br. Mus. (Nat. Hist.) Exp. South-West Arabia 1937-8, 1 (31)).

 

biglumis (Linnaeus)

Vespa biglumis Linnaeus, 1758, Syst. Nat.: 573.

Vespa rupestris Linnaeus, 1758, Syst. Nat.: 573.

Vespa bimaculata Geoffroy, 1785, in Fourcroy, Entomol. Paris 2: 433.

Polistes Geoffroyi Lepeletier and Serville, 1825, Encyc. Méthod. 10: 173.

Polistes dubia Kohl, 1898, Ann. Naturh. Hofmus., Wien 13: 90, non de Saussure, 1867.

Polistes Kohli Dalla Torre, 1904, Genera Insect. 19: 70.  Replacement name for Polistes dubia Kohl, 1898, non de Saussure, 1867.

V. bimacula [!] gFourcroyh; Dalla Torre, 1904, Genera Insect. 19: 72.

Polistes bimaculatus var. Arduinoi Guiglia, 1948, Mem. Soc. Entomol. Ital. 27, Fasc. Suppl.: 14 (key), 22.

gPolistes pamirensis Soikah Zirngiebl, 1955, Mitt. München. Entomol. Ges. 44/45: 381.

Polistes pamirensis var. soikai Zirngiebl, 1955, Mitt. München. Entomol. Ges. 44/45: 383.

Polistes pamirensis var. interruptus Zirngiebl, 1955, Mitt. München. Entomol. Ges. 44/45: 383.

Polistes bimaculatus var. nigrinotum Zirngiebl, 1955, Mitt. München. Entomol. Ges. 44/45: 385.

Polistes biglumis alpium Blüthgen, 1957, Rev. Fac. Sci. Univ. Istanbul, Ser. B, 22 (3): 163.

Polistes geoffiroyi [!]; Das and Gupta, 1984, Orient. Insect. 17: 404.

Distribution: Europe; northern Africa; Turkey; Iran; Mongolia (Guiglia, 1972, Guêpes Soc. Europe Occident. Septentrion.; Giordani Soika, 1976, Acta Zool. Acad. Sci. Hung. 22; Guiglia, 1979, Boll. Mus. Civ. Stor. Nat. Venezia 30; Day, 1979, Biol. J. Linn. Soc. 12).

 

bischoffi Weyrauch

Polistes Bischoffi Weyrauch, 1937, Zool. Jahrb. (Abt. Syst. Ökol. Geogr. Tiere) 70: 274.

Distribution: central and southern Europe; Malta; Turkey (Guiglia, 1972, Guêpes Soc. Europe Occident. Septentrion).

 

chinensis antennalis Pérez

Polistes biglumis var. antennalis Pérez, 1905, Bull. Mus. Hist. Nat. Paris 2: 81.

Polistes insepultus Giordani Soika, 1976, Ann. Hist.-Nat. Mus. Natl. Hung. 68: 288.

Distribution: Mongolia; Korea; Japan (Giordani Soika, 1976, Acta Zool. Acad. Sci. Hung. 22; Kim, 1980, Distr. Atlas Insect. Korea, Ser. 3, Hym. Dipt.; Lee, 1982, Econ. Insect. Fauna China, Fasc. 30).

 

chinensis chinensis (Fabricius)

Vespa chinensis Fabricius, 1793, Entomol. Syst. 2: 261.

Polistes caspicus Yoshikawa, 1962, J. Biol., Osaka City Univ. 13: 21, nomen nudum.

Distribution: Russia: Siberia; China; Hong Kong; Taiwan; Japan: Okinawa; introduced New Zealand (Liu, 1937, Peking Nat. Hist. Bull. 11 (3); Clapperton et al., 1989, New Zealand J. Zool. 16; Starr, 1992, Bull. Natl. Mus. Nat. Sci. 3).

 

dominula (Christ)

Vespa dominula Christ, 1791, Naturgesch. Insect.: 229.

Polistes italica Herrich-Schäffer, 1840, Nomencl. Entomol. 2: 196, nomen nudum.

Polistes pectoralis gMeg.h Herrich-Schäffer, 1841, Fauna Insect. German. 179: 39, 179.6.

Polistes gallica var. Lefebvrei Guérin, 1844, Icon. Règne Anim. 3: 447.

Polistes Bucharensis Erichson, 1849, Mem. Acad. Sci. St. Petersburg (6) 6: 307.

Polistes Lefebvrii [!]; de Saussure, 1853, Ét. Fam. Vesp. 2: 49.

Polistes maculatus Rudow, 1889, Soc. Entomol. 3: 171.

Polistes Merceti Dusmet, 1903, Mem. Soc. Espan. 2 (3): 146 (key), 149.

Polistes gallicus var. rufescens du Buysson, 1912, Ann. Hist. Nat. Délégation Perse II. Entomol. 1: 79.

Polistes gallica var. ornata Weyrauch, 1938, Arbeit. Physiol. Angewand. Entomol. Berlin-Dahlem 5: 278, non Lepeletier, 1836.

Polistes gallica var. pacifica Weyrauch, 1939, Arch. Naturgesch. (N. F.) 8: 155, non Fabricius, 1804.

Polistes gallicus pseudopacificus Giordani Soika, 1970, Ann. Hist.-Nat. Mus. Natl. Hung., Zool. 62: 326.  Replacement name for pacificus Weyrauch, 1939, non Fabricius, 1804.

Polistes gallicus muchei Gusenleitner, 1976, Nachrichtenbl. Bayer. Entomol. 25 (6): 118.

Vespa dominuta [!]; Das and Gupta, 1984, Orient. Insect. 17: 404.

Distribution: central and southern Europe; Turkey; northern Africa; Israel; Syria; Afghanistan; Russia; Iran; Uzbekistan; Turkmenistan; Pakistan; India: Jammu and Kashmir; Mongolia; China; introduced Australia: Western Australia; Chile; Argentina; U.S.A.: transcontinental (Liu, 1937, Peking Nat. Hist. Bull. 11 (3): 313; Guiglia, 1972, Guêpes Soc. Europe Occident. Septentrion.; Day, 1979, Biol. J. Linn. Soc. 12; Hathaway, 1981, Psyche 88; Das and Gupta, 1984, Orient. Insect. 17; Cardale, 1985, in Walton, Zool. Cat. Australia 2; Yildrim and Özbek, 1993, Türk. Entomol. Derg. 17).

 

gallicus (Linnaeus)

Vespa gallica Linnaeus, 1767, Syst. Nat. Ed. 12, 1 (2): 949.

Polistes pictior Radoszkowski, 1872, Horae Entomol. Soc. Ross. 8 (4): v, nomen nudum.

Polistes foederata Kohl, 1898, Ann. Naturh. Hofmus., Wien 13: 90.

Polistes gallicus var. mongolicus du Buysson, 1911, Bull. Mus. Natl. Hist. Nat. 76: 218.

Polistes foederata var. obscuricornis Mader, 1936, Entomol. Zeitschr. 50 (23): 263.

Polistula omissa Weyrauch, 1938, Arbeit. Physiol. Angewand. Entomol. Berlin-Dahlem 5: 277.

Polistes omissus var. ordubadensis Zirngiebl, 1955, Mitt. München Entomol. Ges. 44/45: 381.

Polistes gallicus var. monogolicus [!]; Yoshikawa, 1962, J. Biol. Osaka City Univ. 13: 28.

Polistes omissus kaszabi Giordani Soika, 1970, Ann. Hist.-Nat. Mus. Natl. Hung., Zool. 62: 327.

Polistes foederatus albellus Giordani Soika, 1976, Acta Zool. Acad. Sci. Hung. 22: 272.

Distribution: southern Europe; Ukraine; Russia; Armenia; Azerbaijan; Turkmenistan; Mongolia; China; Iran; Afghanistan; Turkey; Israel; northern Africa; Ethiopia (Zimmermann, 1930, Mitt. Zool. Mus. Berlin 15; Liu, 1937, Peking Nat. Hist. Bull. 11 (3): 313; Guiglia, 1972, Guêpes Soc. Europe Occident. Septentrion.; Gusenleitner, 1985, Nachrichtenbl. Bayer. Entomol. 34 (4); Gusenleitner, 1991, Linzer Biol. Beitr. 23 (2)).

 

indicus Stolfa

Polistes indicus Stolfa, 1934, Bol. Soc. Veneziana Stor. Nat. 1: 47.

Distribution: Afghanistan; Iraq; Iran; Pakistan; United Arab Emirates; Oman (Richards, 1985, Fauna Saudi Arabia 6; Gusenleitner, 2004, Linz. Biol. Beitr. 36 (1)).

 

iranus Guiglia

Polistes iranus Guiglia, 1976, Ann. Mus. Civ. Stor. Nat. Genova 81: 198, nomen nudum.

Polistes iranus Guiglia, 1977 (1976), Boll. Mus. Civ. Stor. Nat. Venezia 28: 99.

Distribution: Iran.

 

loveridgei Bequaert

Polistes loveridgei Bequaert, 1938, Rev. Zool. Bot. Afr. 31: 135.

Distribution: Tanzania; Ethiopia (Giordani Soika, 1952, Riv. Biol. Colon. 11).

 

macrocephalus Bequaert

Polistes macrocephalus Bequaert, 1918, Bull. Am. Mus. Nat. Hist. 39: 261.

Distribution: Congo.

 

marginalis baidoensis Giordani Soika

Polistes marginalis var. baidoensis Giordani Soika, 1944, Atti Inst. Ven. Sci. Lett. Arti, Part II Cl. Sci. Mat. Nat. 103: 177, female, male - Iscia Baidoa, Somalia (lectotype female Venezia).

Distribution: Ethiopia; Somalia.

 

marginalis lindensis Giordani Soika

Polistes marginalis lindensis Giordani Soika, 1981, Boll. Soc. Entomol. Ital. 113: 174.

Distribution: Tanzania.

 

marginalis marginalis (Fabricius)

Vespa marginalis Fabricius, 1775, Syst. Entomol.: 367.

Polistes marginalis arabicus Giordani Soika, 1957, Br. Mus. (Nat. Hist.) Exp. South-West Arabia 1937-8, 1 (31): 482, female, male - Dhala, W. Aden, 4800-5500 piedi (holotype female London).

Distribution: Saudi Arabia; Yemen; Somalia; Ethiopia; Tanzania; Burkina Faso; South Africa: Northern Transvaal, Orange Free State, Kwazulu-Natal, Western and Eastern Cape Prov.; Lesotho (Bequaert, 1938, Rev. Zool. Bot. Afr. 31: 142; Giordani Soika, 1952, Riv. Biol. Colon. 11; Giordani Soika, 1961, South Afr. Anim. Life 8; Das and Gupta, 1984, Orient. Insect. 17; Richards, 1985, Fauna Saudi Arabia 6; Gusenleitner, 2005, Linz. Biol. Beitr. 37 (1)).

 

marginalis meruensis Giordani Soika

Polistes marginalis meruensis Giordani Soika, 1981, Boll. Soc. Entomol. Ital. 113: 173.

Distribution: Kenya.

 

mysteriosus Bequaert

Polistes mysteriosus Bequaert, 1938, Rev. Zool. Bot. Afr. 31: 143.

Distribution: Congo.

 

nimpha (Christ)

Vespa nimpha Christ, 1791, Naturgesch. Insect.: 232.

Vespa diadema Latreille, 1802, Ann. Mus. Hist. Nat. 1: 292, non Christ, 1791.

Polistes opinabilis Kohl, 1898, Ann. Naturh. Hofmus., Wien 13: 90.

Polistes nympha [!]; Weyrauch, 1939, Arch. Naturgesch. (N. F.) 8: 145.

Polistes nimpha var. Moltonii Guiglia, 1944, Atti Soc. Ital. Sci. Nat. 83: 166.

Polistes nimpha var. molotnii [!]; Yoshikawa, 1962, J. Biol. Osaka City Univ. 13: 33.

Polistes nimpha irakensis Gusenleitner, 1976, Nachrichtenbl. Bayer. Entomol. 25 (6): 119.

Distribution: central and southern Europe; Madeiras; northern Africa; Israel; Turkey; Iraq; Azerbaijan; Iran; Pakistan; India: Jammu and Kashmir, Himachal Pradesh; Mongolia; China (Zimmermann, 1930, Mitt. Zool. Mus. Berlin 15; Guiglia, 1972, Guêpes Soc. Europe Occident. Septentrion.; Das and Gupta, 1989, Orient. Insect. 11.; Yildrim and Özbek, 1993, Türk. Entomol. Derg. 17).

 

ornatus Lepeletier

Polistes ornata Lepeletier, 1836, Hist. Nat. Insect. Hym. 1: 531.

Distribution: Liberia; Congo; Kenya (Bequaert, 1938, Rev. Zool. Bot. Afr. 31: 143).

 

riparius Yamane and Yamane

Polistes riparius Yamane and Yamane, 1987, Kontyû 55: 215.

Distribution: Japan.

 

semenowi Morawitz

Polistes semenowi Morawitz, 1889, Horae Entomol. Soc. Ross. 23: 552.

Polistes semenowii [!]; Dalla Torre, 1894, Cat. Hym. 9: 134.

Pseudopolistes semenovi [!]; Weyrauch, 1937, Zool. Jahrb. (Abt. Syst. Ökol. Geogr. Tiere) 70: 265.

Distribution: southern Europe; northern Africa; Turkey; Iran; Turkmenistan (Guiglia, 1972, Guêpes Soc. Europe Occident. Septentrion).

 

sulcifer Zimmermann

Polistes sulcifer Zimmermann, 1930, Mitt. Zool. Mus. Berlin 15: 610.

Polistes sulcifera [!]; Guiglia and Capra, 1933 (1932), Mem. Soc. Entomol. Ital. 11: 166.

Pseudpolistes sulcifer var. similator Zirngiebl, 1955, Mitt. München. Entomol. Ges. 44/45: 384.

Distribution: southern and central Europe; Madeiras; northern Africa; Egypt; Israel; Turkey; Iran (Guiglia, 1972, Guêpes Soc. Europe Occident. Septentrion).

 

tenellus lahejensis Giordani Soika

Polistes tenellus lahejensis Giordani Soika, 1981, Boll. Soc. Entomol. Ital. 113: 175.

Distribution: Yemen.

 

tenellus minutissimus Giordani Soika

Polistes tenellus minutissimus Giordani Soika, 1981, Boll. Soc. Entomol. Ital. 113: 174.

Distribution: Ethiopia.

 

tenellus tenellus du Buysson

Polistes tenellus du Buysson, 1905, Bull. Soc. Entomol. France 1905: 256.

Distribution: Liberia; Nigeria; Cameroon; Equatorial Guinea; Congo Republic; Congo (Bequaert, 1938, Rev. Zool. Bot. Afr. 31: 150).

 

 

SUBGENUS APHANILOPTERUS MEUNIER

 

Aphanilopterus Meunier, 1888, Nat. Sicil. 7: 302, genus.

Type species: Aphanilopterus vagabundus Meunier, 1888 [= Vespa lanio Fabricius, 1775], by monotypy.

Polistoides Dalla Torre, 1904, Genera Insect. 19: 68, name for division II of genus Polistes Latreille in de Saussure, 1854, Ét. Fam. Vesp. 2: 100. Non Polistoides Loew, 1873 (Diptera).

Type species: Polistes subsericeus de Saussure, 1854, by monotypy.

Onerarius Richards, 1973, Rev. Bras. Entomol. 17: 91, 94, 101, subgenus of Polistes Latreille.

Type species: Vespa carnifex Fabricius, 1775, by original designation.

Fuscopolistes Richards, 1973, Rev. Bras. Entomol. 17: 91, 95, 100, subgenus of Polistes Latreille.

Type species: Vespa fuscata Fabricius, 1793, by original designation.

Polistarchus Richards, 1973, Rev. Bras. Entomol. 17: 94, 95, 101, subgenus of Polistes Latreille.

Type species: Vespa canadensis Linnaeus, 1758, by original designation.

Palisotius Richards, 1973, Rev. Bras. Entomol. 17: 95, 96, 100, subgenus of Polistes Latreille.

Type species: Polistes major Palisot de Beauvois, 1818, by original designation.

Epicnemius Richards, 1973, Rev. Bras. Entomol. 17: 96, 102, subgenus of Polistes Latreille.

Type species: Polistes bicolor Lepeletier, 1836, by original designation.

 

actaeon Haliday

Polistes actaeon Haliday, 1836, Trans. Linn. Soc. London 17: 323.

Polistes acteon [!]; de Saussure, 1854, Ét. Fam. Vesp. 2: 98.

Polistes limai von Ihering, 1903, Ann. Soc. Entomol. France 72: 145.

Distribution: Brazil; Paraguay; Argentina: Misiones (Richards, 1978, Soc. Wasps Am.).

 

adelphus Richards

Polistes adelphus Richards, 1978, Soc. Wasps. Am.: 501.

Distribution: Colombia; Ecuador.

 

angulinus Richards

Polistes erythrogaster var. angulinus Richards, 1951, in Richards and Richards, Trans. R. Entomol. Soc. London 102: 92.

Distribution: Guyana.

 

annularis (Linnaeus)

Vespa annularis Linnaeus, 1763, Amoenitat. Acad. 6: 413.

Vespa cincta Drury, 1773, Illus. Nat. Hist. 2: index to vol. 1.

Vespa virginiensis [!]; Dalla Torre, 1894, Cat. Hym. 9: 122.

Vespa annulata [!]; Dalla Torre, 1894, Cat. Hym. 9: 123.

Polistes annuralis [!]; Yamane, 1969, J. Fac. Sci., Hokkaido Univ., Ser. VI, Zool. 17: 79.

Distribution: U.S.A.: Massachusetts to Florida, west to South Dakota, south to Texas (Richards, 1978, Soc. Wasps. Am.).

 

apachus de Saussure

Polistes apachus de Saussure, 1857, Ann. Soc. Entomol. France (3) 5: 314.

Polistes texanus Cresson, 1872, Trans. Am. Entomol. Soc. 4: 246.

Distribution: U.S.A.: Kansas to Colorado, south to Texas, occasionally Iowa, California, Florida; Mexico: Baja California, Nuevo León, San Luis Potosí, Coahuila, Sonora, Durango (Richards, 1978, Soc. Wasps. Am.; Krombein, 1979, in Krombein et al., Cat. Hym. Am. N. of Mexico).

 

apicalis de Saussure

Polistes apicalis de Saussure, 1858, Rev. Mag. Zool. (2) 10: 260.

Distribution: Mexico: Veracruz, Chiapas; Belize; Guatemala; Honduras; Costa Rica; French Guiana (Richards, 1978, Soc. Wasps Am.; West-Eberhard et al., 1995, in Hanson and Gauld, Hym. Costa Rica).

 

arizonensis Snelling

Polistes exclamans arizonensis Snelling, 1954, J. Kans. Entomol. Soc. 27: 152.

Distribution: U.S.A.: Arizona, California; Mexico: Sonora (Krombein et al., 1958, Hym. Am. N. of Mexico Suppl.).

 

aterrimus de Saussure

Polistes aterrimus de Saussure, 1853, Ét. Fam. Vesp. 2: 73.

Distribution: Costa Rica; Colombia; Ecuador; Peru; Venezuela; Bolivia (Richards, 1978, Soc. Wasps Am.; West-Eberhard et al., 1995, in Hanson and Gauld, Hym. Costa Rica).

 

atrox Richards

Polistes atrox Richards, 1978, Soc. Wasps. Am.: 542.

Distribution: Brazil: Pará.

 

aurifer de Saussure

Polistes aurifer de Saussure, 1853, Ét. Fam. Vesp. 2: 78.

Polistes anaheimensis Provancher, 1888, Addit. Corr. Faune Entomol. Can., Hym.: 423.

Polistes fuscatus var. utahensis Hayward, 1933, Proc. Utah Acad. Sci. 10: 142.

Polistes fuscatus var. centralis Hayward, 1933, Proc. Utah Acad. Sci. 10: 143.

Polistes fuscatus var. (or subsp.) connectens Bequaert, 1940, J. New York Entomol. Soc. 48: 21.

Polistes fuscatus var. (or subsp.) montanus Bequaert, 1940, J. New York Entomol. Soc. 48: 26.

Distribution: Canada: Alberta, British Columbia; U. S .A.: Montana to Washington, south to California, Arizona, New Mexico, Texas; Mexico: Baja California, Sonora, Chihuahua, Coahuila; introduced U.S.A.: Hawaii, Society Is.: Bora Bora, Raiatea, Tahiti (Richards, 1978, Soc. Wasps. Am.; MacLean et al., 1978, Great Lakes Entomol. 11; Krombein, 1979, in Krombein et al., Cat. Hym. Am. N. of Mexico).

 

bahamensis Bequaert and Salt

Polistes bahamensis Bequaert and Salt, 1931, Ann. Entomol. Soc. Am. 24: 793.

Polistes bahamensis var. bilineolatus Bequaert and Salt, 1931, Ann. Entomol. Soc. Am. 24: 796.

Polistes bahamensis var. picturatus Bequaert and Salt, 1931, Ann. Entomol. Soc. Am. 24: 797.

Polistes exclamans var. (or subsp.) louisianus Bequaert, 1940, J. New York Entomol. Soc. 48: 14.

Distribution: U.S.A.: Florida to Louisiana; Bahamas (Krombein, 1979, in Krombein et al., Cat. Hym. Am. N. of Mexico; Snelling, 1983, Pan-Pacif. Entomol. 59).

 

bellicosus Cresson

Polistes bellicosus Cresson, 1872, Trans. Am. Entomol. Soc. 4: 247.

Distribution: North Carolina to Florida, west to Texas (Richards, 1978, Soc. Wasps. Am.).

 

bequaertellus Snelling

Polistes bequaertellus Snelling, 1983, Pan-Pacif. Entomol. 59: 269.

Distribution: El Salvador.

 

bequaertianus Willink

Polistes bequaertiana Willink, 1954, Acta Zool. Lilloana 14: 255.

Distribution: Argentina: Misiones.

 

bicolor Lepeletier

Polistes bicolor Lepeletier, 1836, Hist. Nat. Insect. Hym. 1: 521.

Polistes deuteroleucus Ducke, 1906, Rev. Entomol., Caen 25: 10.

Polistes bicolor var unicolor Ducke, 1908, Deutsch. Entomol. Zeitschr. 6: 700, non Lepeletier, 1836.

Polistes sosia Bequaert, 1937, Arch. Inst. Biol. Veg. 3: 190.

Polistes duckei Bequaert, 1937, Arch. Inst. Biol. Veg. 3: 192.  Replacement name for bicolor var. unicolor Ducke, 1908, non Lepeletier, 1836.

Distribution: Costa Rica; Panama; Colombia; Ecuador; Peru; Venezuela; Suriname; French Guiana; Brazil: Amapá, Pará, Amazonas, Acre, Mato Grosso, Minas Gerais (Richards, 1978, Soc. Wasps Am.; West-Eberhard et al., 1995, in Hanson and Gauld, Hym. Costa Rica).

 

biguttatus Haliday

Polistes biguttatus Haliday, 1836, Trans. Linn. Soc. London 17: 323.

? Polistes bipustulatus [!]; Smith, 1875, Entomol. Mon. Mag. 12: 156.

Polistes bipustulstus [!]; Yoshikawa, 1962, J. Biol., Osaka City Univ. 13: 22.

Distribution: Brazil: Rio de Janeiro to Rio Grande do Sul (Richards, 1978, Soc. Wasps Am.).

 

billardieri biglumoides Ducke

Polistes biglumoides Ducke, 1904, Bol. Mus. Goeldi, Pará 4: 369.

Distribution: Colombia; Peru; Venezuela; Brazil: Amapá, Pernambuco, to São Paulo, Matto Grosso; Bolivia (Richards, 1978, Soc. Wasps Am.).

 

billardieri billardieri Fabricius

Polistes billardieri Fabricius, 1804, Syst. Piez.: 274.

Polistes ruficornis var. demeraraensis Bequaert, 1937, Arch. Inst. Biol. Veg. 3: 181.

Distribution: Colombia; Guyana; Suriname; Brazil: Roraima, Amazonas, Mato Grosso (Richards, 1978, Soc. Wasps Am.).

 

billardieri ruficornis de Saussure

Polistes ruficornis de Saussure, 1853, Ét. Fam. Vesp. 2: 85.

Distribution: Brazil: Goiás, Minas Gerais, São Paulo, Mato Grosso, Paraná, Rio Grande do Sul; Paraguay; Uruguay; Argentina: Misiones to Salta; Bolivia (Richards, 1978, Soc. Wasps Am.).

 

binotatus de Saussure

Polistes binotatus de Saussure, 1853, Ét. Fam. Vesp. 2: 87.

Distribution: Brazil: Rio de Janeiro (Richards, 1978, Soc. Wasps Am.).

 

boharti Snelling

Polistes boharti Snelling, 1983, Pan-Pacif. Entomol. 59: 272.

Distribution: Mexico: Sinaloa, Nayarit, Guerrero, Colima, Oaxaca, Veracruz.

 

brevifissus Richards

Polistes brevifissus Richards, 1978, Soc. Wasps Am.: 481.

Distribution: Brazil: Amapá, Roraima, Pará, Mato Grosso, Bahia to Rio Grande do Sul; Argentina: Buenos Aires.

 

buyssoni Brèthes

Polistes cavapyta var. thoracica du Buysson, 1906, Entomol. Medd. 3: 19, non Fox, 1898.

Polistes cavapyta var. buyssoni Brèthes, 1909, An. Mus. Nac. Hist. Nat. Buenos Aires (3) 17: 456.  Replacement name for cavapyta var. thoracica du Buysson, 1906, non Fox, 1898.

Distribution: Bolivia; Argentina: Jujuy to Rio Negro; introduced Chile (Willink, 1952, Acta Zool. Lilloana 10; Pérez-D'Angello, 1970, Not. Mens. Mus. Nac. Hist. Nat. 15 (169); Willink and Chiappa, 1993, Acta Entomol. Chil. 18).

 

canadensis borientalis Richards

Polistes canadensis borientalis Richards, 1978, Soc. Wasps Am.: 494.

Distribution: Brazil: Rio Grande do Norte, Paraiba, Pernambuco, Bahia.

 

canadensis canadensis (Linnaeus)

Vespa canadensis Linnaeus, 1758, Syst. Nat.: 574.

gVespa crabro marribous Linn.h Christ, 1791, Naturgesch. Insect.: 217.

? Polistes urceolata gKlugh Erichson, 1848, in Schomburgk, Reisen Br.-Guiana III: 590.

? Polistes urceolicus [!]; Dalla Torre, 1904, Genera Insect. 19: 71.

Polistes unicolor Lepeletier, 1836, Hist. Nat. Insect. Hym. 1: 520.

Polistes panamensis Holmgren, 1868, K. Svenska Fregat. Eugenies Resa Omkring Jorden, Zool. 1, Insect., Hym.: 439.

Polistes canadensiss [!]; Yoshikawa, 1962, J. Biol. Osaka City Univ. 13: 26.

Polistes canadiensis [!]; Maes, 1988, Rev. Nicarag. Entomol. 5: 39.

Distribution: U.S.A.: Arizona; Mexico: Sonora, Tamaulipas, Nayarit to Oaxaca; Guatemala; El Salvador; Honduras; Costa Rica; Panama; Colombia; Peru; Venezuela; Guyana; Suriname; French Guiana; Brazil: Amazonas to Pará, Bahia and Mato Grosso south to São Paulo; Bolivia; Paraguay; Argentina: Corrientes, Misiones, Tucumán (Bequaert, 1944, Bull. Mus. Comp. Zool. 94; Richards, 1978, Soc. Wasps Am.).

 

candidoi von Ihering

Polistes candidoi von Ihering, 1903, Ann. Soc Entomol. France 72: 144.

Distribution: Bolivia.

 

carnifex boliviensis Bequaert

Polistes carnifex var. boliviensis Bequaert, 1936, Rev. Entomol., Rio de Janeiro 6: 381.

Distribution: Colombia; Peru; Bolivia (Richards, 1978, Soc. Wasps Am.).

 

carnifex carnifex (Fabricius)

Vespa carnifex Fabricius, 1775, Syst. Entomol.: 365.

? Polistes variegata Lepeletier, 1836, Hist. Nat. Insect. Hym. 1: 523.

Polistes onerata Lepeletier, 1836, Hist. Nat. Insect. Hym. 1: 524.

Polistes valida Say, 1837, Boston J. Nat. Hist. 1: 389 - gMexicoh (destroyed).

Polistes transverso-strigata Spinola, 1851, preprint of Mem. R. Accad. Sci. Torino (2) 13: 62.

Polistes transversosignatus [!]; de Saussure, 1853, Ét. Fam. Vesp. 2: 90.

Polistes transversesignata [!]; Schulz, 1906, Spolia Hym.: 226.

Polistes transversignatus [!]; Yoshikawa, 1962, J. Biol. Osaka City Univ. 13: 23.

Distribution: U.S.A.: Arizona, Texas; Mexico; Belize; Guatemala; Honduras; Nicaragua; Costa Rica; Panama; Colombia; Ecuador; Peru; Brazil: Pernambuco to Amazonas, south to Rio Grande do Sul; Guyana; Suriname; Paraguay; Argentina: Misiones; introduced U.S.A.: Hawaii (Richards, 1978, Soc. Wasps Am.; Krombein, 1979, in Krombein et al., Cat. Hym. Am. N. of Mexico; Nishida, 1992, Hawaiian Terr. Arth. Checklist; West-Eberhard et al., 1995, in Hanson and Gauld, Hym. Costa Rica).

 

carnifex rufipennis Latreille

Polistes rufipennis Latreille, 1817, in Humboldt and Bonpland, Voyage, Zool. 2, Insect. 2: 97.

Polistes chlorostoma Lepeletier, 1836, Hist. Nat. Insect. Hym. 1: 521.

Polistes transverso-strigata var. ochreata gKL.h Spinola, 1851, Mem. R. Accad. Sci. Torino (2) 13: 63.

Distribution: Honduras; Colombia; Trinidad; Tobago; Suriname; French Guiana (Bequaert, 1944, Bull. Mus. Comp. Zool. 94; Richards, 1978, Soc. Wasps Am.).

 

carolina (Linnaeus)

Vespa carolina Linnaeus, 1767, Syst. Nat., Ed. 12, 1 (2): 948.

Vespa nigripennis DeGeer, 1773, Mem. Hist. Insect. 3: 582, pl. 29 fig. 10.

Polistes rubiginosa Lepeletier, 1836, Hist. Nat. Insect. Hym. 1: 524.

Polistes ruginosus [!]; Gowdey, 1926, Entomol. Bull., Dept. Agric. Jamaica 4: 95.

Polistes rubigenosus [!]; Yoshikawa, 1962, J. Biol. Osaka City Univ. 13: 35.

Distribution: U.S.A.: New York to Florida, west to Kansas and Texas; introduced Bermuda (Richards, 1978, Soc. Wasps. Am.; Krombein, 1979, in Krombein et al., Cat. Hym. Am. N. of Mexico).

 

cavapyta de Saussure

Polistes cavapyta de Saussure, 1853, Ét. Fam. Vesp. 2: 87.

Distribution: Brazil: Rio Grande do Sul; Paraguay; Uruguay; Argentina: Misiones to Jujuy, south to Buenos Aires (Willink, 1952, Acta Zool. Lilloana 10; Willink and Chiappa, 1993, Acta Entomol. Chil. 18).

 

cavapytiformis Richards

Polistes cavapytiformis Richards, 1978, Soc. Wasps Am.: 481.

Distribution: Brazil: Paraná to Rio Grande do Sul.

 

cinerascens de Saussure

Polistes cinerascens de Saussure, 1854, Ét. Fam. Vesp. 2: 99.

Polistes orbitalis Kirby, 1888, Ann. Mag. Nat. Hist. (6) 2: 143.

Distribution: Ecuador; Brazil: Bahia to Mato Grosso and Santa Catarina; Bolivia; Paraguay; Uruguay; Argentina: Misiones and Jujuy to Buenos Aires (Richards, 1978, Soc. Wasps Am.).

 

claripennis Ducke

Polistes claripennis Ducke, 1904, Bol. Mus. Goeldi, Pará 4: 370.

Polistes synoecoides Ducke, 1907, Bol. Mus. Goeldi, Pará 5: 196.

Distribution: Brazil: Pará, Amazonas; Peru (Richards, 1978, Soc. Wasps Am.).

 

comanchus comanchus de Saussure

Polistes comanchus de Saussure, 1857, Ann. Soc. Entomol. France (3) 5: 314.

Polistes commanchus [!]; Richards, 1978, Soc. Wasps Am.: 485.

Distribution: U.S.A.: Texas, New Mexico; Mexico: Coahuila, Durango (Richards, 1978, Soc. Wasps. Am.; Krombein, 1979, in Krombein et al., Cat. Hym. Am. N. of Mexico).

 

comanchus navajoe Cresson

Polistes navajoe Cresson, 1868, Trans. Am. Entomol. Soc. 1: 383.

Polistes novajoe [!]; Dalla Torre, 1894, Cat. Hym. 9: 132.

Polistes novajae [!]; Dalla Torre, 1894, Cat. Hym. 9: 177.

Distribution: U.S.A.: Texas to Arizona, California; Mexico: Sonora, Chihuahua, Durango, Sinaloa (Richards, 1978, Soc. Wasps. Am.: 485; Krombein, 1979, in Krombein et al., Cat. Hym. Am. N. of Mexico).

 

consobrinus de Saussure

Polistes consobrinus de Saussure, 1858, Rev. Mag. Zool. (2) 10: 259.

Distribution: Brazil: Paraná, Rio Grande do Sul; Paraguay; Uruguay; Argentina: Buenos Aires, Correnties, Entre Rios, Santa Fé (Richards, 1978, Soc. Wasps. Am.).

 

crinitus americanus (Fabricius)

Vespa americana Fabricius, 1775, Syst. Entomol.: 370.

Polistes madoci Kirby, 1884, Ann. Mag. Nat. Hist. (5) 13: 411.

Polistes madocii [!]; Yoshikawa, 1962, J. Biol., Osaka Univ. 13: 30.

Distribution: Puerto Rico; Mona Is.; St. Thomas (Richards, 1978, Soc. Wasps Am.).

 

crinitus crinitus (Felton)

Vespa crinita Felton, 1765 (1764), Philos. Trans. R. Soc. London 54: 54.

Vespa tricolor Fabricius, 1775, Syst. Entomol.: 369, non Pallas, 1771.

Polistes media Palisot de Beauvois, 1818, Insect. Recueill. Afrique Amérique, livr. 12: 207.

Distribution: Jamaica; Dominican Republic; Montserrat (Schulz, 1906, Spolia Hym.; Richards, 1978, Soc. Wasps Am.).

 

crinitus multicolor (Olivier)

Vespa multicolor Olivier, 1792, Encyc. Méthod. 6: 691.

? Vespa dominicensis Vallot, 1802, Concord. Syst.: 168.

Polistes crinitus var. insulicola Bequaert and Salt, 1931, Ann. Entomol. Soc. Am. 24: 793.

Distribution: Guadeloupe; St. Croix; St. Barthélemy; St. Kitts; Anguilla; Nevis; Montserrat; Antigua; Dominica (de Saussure, 1854, Ét. Fam. Vesp. 2; Richards, 1978, Soc. Wasps Am.).

 

cubensis Lepeletier

Polistes cubensis Lepeletier, 1836, Hist. Nat. Insect. Hym. 1: 526.

Distribution: Cuba; introduced Venezuela (Richards, 1978, Soc. Wasps Am.).

 

davillae Richards

Polistes davillae Richards, 1978, Soc. Wasps Am.: 540.

Distribution: Brazil: Amazonas, Mato Grosso.

 

deceptor Schulz

Polistes deceptor Schulz, 1905, Hym.-Studien: 119.

Polistes rhodostoma Ducke, 1905, Rev. Entomol., Caen 24: 23.

Distribution: Panama; Ecuador; Peru; Guyana; Suriname; Brazil: Pará, São Paulo; Bolivia; Paraguay; Argentina: Misiones (Richards, 1978, Soc. Wasps Am.).

 

dominicus (Vallot)

Vespa dominica Vallot, 1802, Concord. Syst.: 168.

Polistes cincta Lepeletier, 1836, Hist. Nat. Insect. Hym. 1: 522, non Drury, 1773.

Polistes cinctus barbadensis Richards, 1978, Soc. Wasps Am.: 479.

Distribution: St. Kitts; Montserrat; Martinique; Barbados; Santa Lucia; St. Vincent; Mustique Is.; Carriacou Is.; Trinidad (Snelling, 1983, Pan-Pacif. Entomol. 59).

 

dorsalis californicus Bohart

Polistes hunteri californicus Bohart, 1949, Pan-Pacif. Entomol. 25: 101.

Distribution: U.S.A.: California, Arizona; Mexico: Baja California, Chihuahua, Sonora, Sinaloa (Richards, 1978, Soc. Wasps. Am.).

 

dorsalis clarionensis Bohart

Polistes hunteri clarionensis Bohart, 1949, Pan-Pacif. Entomol. 25: 102.

Distribution: Mexico: Revilla Gigedo group and Clarion Is.

 

dorsalis dorsalis (Fabricius)

Vespa dorsalis Fabricius, 1775, Syst. Entomol.: 367.

Polistes fuscatus (var. or subsp.) hunteri Bequaert, 1940, J. New York Entomol. Soc. 48: 23.

Distribution: U.S.A.: Virginia to Florida, west to Arizona, California; Mexico: San Luis Potosí, Sonora, Nayarit, D. F., Morelos, Guerrero; Jamaica (van der Vecht, 1970, Proc. K. Ned. Akad. Wet (ser. C) 73; Richards, 1978, Soc. Wasps. Am.).

 

dorsalis maritimus Bequaert

Polistes fuscatus var (or subsp.) maritimus Bequaert, 1940, J. New York Entomol. Soc. 48: 25.

Distribution: Bahamas: Andros Is.

 

dorsalis neotropicus Bequaert

Polistes fuscatus var (or subsp.) neotropicus Bequaert, 1940, J. New York Entomol. Soc. 48: 28.

Distribution: U.S.A.: Colorado, New Mexico, Texas; Mexico: Veracruz, Oaxaca; Guatemala; Honduras; Costa Rica (Bohart, 1949, Pan-Pacif. Entomol. 25; van der Vecht, 1970, Proc. K. Ned. Akad. Wet (ser. C) 73; Richards, 1978, Soc. Wasps. Am.).

 

eburneus Bequaert

Polistes canadensis var. (or subsp.) eburneus Bequaert, 1943, Bol. Entomol. Venezolana 2: 118.

Distribution: Paraguay.

 

erythrocephalus Latreille

Polistes erythrocephala Latreille, 1813, in Humboldt and Bonpland, Voyage, Zool. 2, Insect. 2: 96.

Distribution: Nicaragua; Costa Rica; Panama; Colombia; Ecuador; Peru; Bolivia; Brazil: Pará to Acre, south to Paraná (Richards, 1978, Soc. Wasps Am.).

 

exclamans Viereck

Polistes exclamans Viereck, 1906, Trans. Am. Entomol. Soc. 32: 201.

Polistes exclamans durangoensis Snelling, 1955, Am. Mus. Novitates 1701: 7.

Polistes instabilis coahuilae Richards, 1978, Soc. Wasps Am.: 510.

Distribution: U.S.A.: New Jersey to Florida, west to Nebraska, Colorado, Texas; Mexico: Chihuahua to Jalisco, Hidalgo; introduced U.S.A.: Hawaii (Krombein, 1979, in Krombein et al., Cat. Hym. Am. N. of Mexico; Snelling, 1983, Pan-Pacif. Entomol. 59; Nishida, 1992, Hawaiian Terr. Arth. Checklist).

 

ferreri de Saussure

Polistes ferreri de Saussure, 1853, Ét. Fam. Vesp. 2: 77.

Distribution: Brazil: Bahia to Rio Grande do Sul, Mato Grosso; Bolivia; Uruguay; Argentina: Buenos Aires, Tucumán (Richards, 1978, Soc. Wasps Am.).

 

flavus Cresson

Polistes flavus Cresson, 1868, Trans. Am. Entomol. Soc. 1: 383.

Distribution: U.S.A.: California (southern), Nevada to Colorado, Arizona to Texas (western); Mexico: Sonora, Chihuahua, Coahuila (Richards, 1978, Soc. Wasps. Am.; Krombein, 1979, in Krombein et al., Cat. Hym. Am. N. of Mexico).

 

franciscanus Richards

Polistes franciscanus Richards, 1978, Soc. Wasps Am.: 521.

Distribution: Guatemala.

 

fuscatus (Fabricius)

? Vespa tibialis Olivier, 1792, Encyc. Méthod. Insect. 6: 690, non Gmelin, 1790.

Vespa fuscata Fabricius, 1793, Entomol. Syst. 2: 260.

Vespa nestor Fabricius, 1798, Suppl. Entomol. Syst.: 262.

Polistes pallipes Lepeletier, 1836, Hist. Nat. Insect. Hym. 1: 530.

Polistes exilis de Saussure, 1853, Ét. Fam. Vesp. 2: 85.

Polistes variatus Cresson, 1872, Trans. Am. Entomol. Soc. 4: 247.

Polistes fortunatus Kirby, 1884, Ann. Mag. Nat. Hist. (5) 13: 410.

Polistes pallidipes Dalla Torre, 1894, Cat. Hym. 9: 133, unnecessary emendation.

Polistes fuscatus vtriatus [!]; Hayward, 1933, Proc. Utah Acad. Sci. 10: 142.

Polistes fuscatus var. (or subsp.) laurentianus Bequaert, 1942, Can. Entomol. 74: 159.

Polistes varistus [!]; Yoshikawa, 1962, J. Biol. Osaka City Univ. 13: 27.

Distribution: Canada; U.S.A.: Maine south to Florida, west to Nebraska, south to Texas; introduced Bermuda; Jamaica; Barbados; Cape Verde Is.; Ascension Is. (Richards, 1978, Soc. Wasps. Am.; MacLean et al., 1978, Great Lakes Entomol. 11; Krombein, 1979, in Krombein et al., Cat. Hym. Am. N. of Mexico).

 

geminatus geminatus Fox

Polistes geminatus Fox, 1898, Proc. Acad. Nat. Sci. Philadelphia 1898: 448.

Polistes gaminatus [!]; Yoshikawa, 1962, J. Biol. Osaka City Univ. 13: 28.

Distribution: French Guiana; Brazil: Pará, Minas Gerais, Goiás, Mato Grosso, São Paulo (Richards, 1978, Soc. Wasps Am.).

 

geminatus guyanensis Cameron

Polistes guyanensis Cameron, 1912, Timehri 2: 221.

Distribution: Guyana.

 

goeldii Ducke

Polistes goeldii Ducke, 1904, Bol. Mus. Goeldi, Pará 4: 368.

Distribution: El Salvador; Costa Rica; Colombia; Trinidad; Ecuador; Peru; Brazil: Pará, Amazonas, Mato Grosso (Richards, 1978, Soc. Wasps Am.; West-Eberhard et al., 1995, in Hanson and Gauld, Hym. Costa Rica).

 

huacapistana Richards

Polistes huacapistana Richards, 1978, Soc. Wasps Am.: 496.

Distribution: Peru.

 

incertus Cresson

Polistes incertus Cresson, 1865, Proc. Entomol. Soc. Philadelphia 4: 166.

Distribution: Cuba, Is. de la Juventud; Haiti; Dominican Republic; Tortuga Is. (Richards, 1978, Soc. Wasps Am.).

 

infuscatus anduzei Bequaert

Polistes canadensis var. (or subsp.) anduzei Bequaert, 1943, Bol. Entomol. Venezolana 2: 114.

Distribution: Venezuela; Peru (Richards, 1978, Soc. Wasps Am.).

 

infuscatus ecuadorius Richards

Polistes infuscatus ecuadorius Richards, 1978, Soc. Wasps Am.: 484.

Distribution: Ecuador; Colombia.

 

infuscatus infuscatus Lepeletier

Polistes infuscata Lepeletier, 1836, Hist. Nat. Insect. Hym. 1: 520.

Polistes canadensis amazonicus Schulz, 1905, Hym.-Studien: 113.

Distribution: Costa Rica; Panama; Colombia; Venezuela; Guyana; Suriname; French Guiana; Ecuador; Peru; Brazil: Pará (Schulz, 1905, Hym.-Studien; Bequaert, 1944, Bull. Mus. Comp. Zool. 94; Richards, 1978, Soc. Wasps Am.; West-Eberhard et al., 1995, in Hanson and Gauld, Hym. Costa Rica).

 

infuscatus mariae Richards

Polistes infuscatus mariae Richards, 1978, Soc. Wasps Am.: 484.

Distribution: Colombia.

 

infuscatus metensis Richards

Polistes infuscatus metensis Richards, 1978, Soc. Wasps Am.: 484.

Distribution: Colombia.

 

instabilis de Saussure

Polistes instabilis de Saussure, 1853, Ét. Fam. Vesp. 2: 91, pl. X fig. 2, pl. XI fig. 1.

Polistes instbailis [!]; Yoshikawa, 1962, J. Biol., Osaka Univ. 13: 24.

Distribution: Mexico: Sonora and Tamaulipas to Yucatán; Belize; Guatemala; Honduras; Nicaragua; Costa Rica; ? introduced U.S.A.: southern Texas (Snelling, 1974, Proc. Entomol. Soc. Washington 76; Richards, 1978, Soc. Wasps Am.).

 

kaibabensis Hayward

Polistes canadensis var. kaibabensis Hayward, 1932, Proc. Utah Acad. Sci. 9: 89.

Distribution: U.S.A.: Arizona (Grand Canyon), southern Utah (Krombein, 1979, in Krombein et al., Cat. Hym. Am. N. of Mexico).

 

lanio lanio (Fabricius)

Vespa lanio Fabricius, 1775, Syst. Entomol.: 365.

Aphanilopterus vagabundus Meunier, 1888, Nat. Sicil. 7: 302.

Polistes canadensis trinitatis Giordani Soika, 1965, Boll. Entomol. Soc. Ital. 95: 27, non Bequaert, 1937.

Polistes lanio soikai Richards, 1978, Soc. Wasps Am.: 490, non Zirngiebl, 1955.  Replacement name for canadensis trinitatis Giordani Soika, 1965, non Bequaert, 1937.

Polistes gefoveatus Richardsh Almeida and Castro, 1991, Acta Biol. Par., Curitiba 20: 50, nomen nudum.

Distribution: Colombia; Peru; Trinidad; Guyana; Brazil: Amapá to Acre, Pará to Rio Grande do Sul; Bolivia; Paraguay; Argentina: Corrientes, Misiones (Richards, 1978, Soc. Wasps Am.).

 

lanio satanulus Bequaert

Polistes canadensis var. (or subsp.) satanulus Bequaert, 1940, Entomol. News 51: 82.

Distribution: Brazil: Santa Catarina.

 

lanio weberi Bequaert

Polistes canadensis var. (or subsp.) weberi Bequaert, 1940, J. New York Entomol. Soc. 48: 12.

Distribution: Venezuela.

 

lineonotus Bohart

Polistes exclamans lineonotus Bohart, 1949, Pan-Pacif. Entomol. 25: 98.

Distribution: Mexico: Baja California (Snelling, 1983, Pan-Pacif. Entomol. 59).

 

major bonaccensis Bequaert

Polistes major var. bonaccensis Bequaert, 1937, Arch. Inst. Biol. Veg. 3: 174.

Distribution: Honduras: Bonacca Is.

 

major castaneicolor Bequaert

Polistes major var. castaneicolor Bequaert, 1936, Entomol. News 47: 12.

Distribution: U.S.A.: southern Arizona, New Mexico; Mexico: Baja California, Sonora, Chihuahua, Jalisco (Richards, 1978, Soc. Wasps Am.; Krombein, 1979, in Krombein et al., Cat. Hym. Am. N. of Mexico).

 

major colombianus Bequaert

Polistes major var. (or subsp.) colombianus Bequaert, 1940, Entomol. News 51: 80.

Distribution: Colombia.

 

major major Palisot de Beauvois

Polistes major Palisot de Beauvois, 1818, Insect. Recueill. Afrique Amérique, livr. 12: 206.

Polistes maior [!]; Schulz, 1906, Spolia Hym.: 227.

Polistes major var. (or subsp.) bakeri Bequaert, 1940, J. New York Entomol. Soc. 48: 15.

Distribution: U.S.A.: southern Arizona, New Mexico, Texas, Florida; Cuba; Bahamas; Navassa Is.; Dominican Republic; Jamaica; Mexico; Guatemala, El Salvador; Honduras; Nicaragua; Costa Rica; Venezuela; Trinidad; Brazil: Pará, Amazonas, Espírito Santo, São Paulo, Mato Grosso; ? introduced Puerto Rico (Richards, 1978, Soc. Wasps Am.; Krombein, 1979, in Krombein et al., Cat. Hym. Am. N. of Mexico; West-Eberhard et al., 1995, in Hanson and Gauld, Hym. Costa Rica).

 

major weyrauchi Bequaert

Polistes major var. (or subsp.) weyrauchi Bequaert, 1940, Entomol. News 51: 79.

Distribution: Panama; Colombia; Venezuela; Peru (Richards, 1978, Soc. Wasps Am.).

 

maranonensis Willink

Polistes versicolor var. flavescens Weyrauch, 1943, Bol. Mus. Hist. Nat. gJavier Pradoh 7: 71, nomen nudum.

Polistes maranonensis Willink, 1964, Acta Zool. Lilloana 20: 16.

Distribution: Peru (Richards, 1978, Soc. Wasps Am.).

 

melanosoma de Saussure

Polistes melanosoma de Saussure, 1853, Ét. Fam. Vesp. 2: 76.

Distribution: Guyana; Brazil: Amazonas, Espírito Santo to Rio Grande do Sul; Paraguay; Argentina: Misiones (Bequaert, 1937, Arch. Inst. Biol. Veg. 3; Bequaert, 1944, Bull. Mus. Comp. Zool. 94; Richards, 1978, Soc. Wasps Am.).

 

melanotus Richards

Polistes melanotus Richards, 1978, Soc. Wasps Am.: 487.

Distribution: Colombia.

 

metricus Say

Polistes metrica Say, 1831, Am. Entomol.: 15.

Distribution: U.S.A.: New York to Florida, west to Nebraska, south to Texas (Krombein, 1979, in Krombein et al., Cat. Hym. Am. N. of Mexico).

 

mexicanus Bequaert

Polistes canadensis var. (or subsp.) mexicanus Bequaert, 1940, J. New York Entomol. Soc. 48: 10.

Polistes canadensis var. (or subsp.) wheeleri Bequaert, 1940, J. New York Entomol. Soc. 48: 12.

Distribution: Mexico: Sonora, Veracruz, Puebla, D. F., Morelos, Guerrero, Oaxaca (Snelling, 1974, Proc. Entomol. Soc. Washington 76; Richards, 1978, Soc. Wasps Am.).

 

minor Palisot de Beauvois

Polistes minor Palisot de Beauvois, 1818, Insect. Recueill. Afrique Amérique, livr. 12: 207.

Polistes hertwigi Schulz, 1903, Sitzungsber. Math.-Phys. Kl. Bayer. Akad. Wiss. 33: 486.

Distribution: Dominican Republic; Haiti; Puerto Rico; Tortuga Is. (Richards, 1978, Soc. Wasps Am.).

 

moraballi Richards

Polistes erythrogaster var. moraballi Richards, 1951, in Richards and Richards, Trans. R. Entomol. Soc. London 102: 92.

Distribution: Guyana.

 

myersi curassavicus Richards

Polistes myersi curassavicus Richards, 1978, Soc. Wasps Am.: 507.

Distribution: Netherlands Antilles: Curaçao; Aruba; Bonaire.

 

myersi myersi Bequaert

Polistes versicolor var. myersi Bequaert, 1934, Rev. Entomol., Rio de Janeiro 4 : 151.

Distribution: Costa Rica; Panama; Colombia; Venezuela (Richards, 1978, Soc. Wasps Am.; West-Eberhard et al., 1995, in Hanson and Gauld, Hym. Costa Rica).

 

niger Brèthes

Polistes niger Brèthes, 1903, An. Mus. Nac. Hist. Nat. Buenos Aires (3) 9: 20.

Distribution: Brazil: São Paulo, Minas Gerais, Mato Grosso (Bequaert, 1937, Arch. Inst. Biol. Veg. 3; Richards, 1978, Soc. Wasps Am.: 551).

 

ninabamba Richards

Polistes ninabamba Richards, 1978, Soc. Wasps Am.: 500.

Distribution: Peru.

 

notatipes Richards

Polistes notatipes Richards, 1978, Soc. Wasps Am: 488.

Distribution: Brazil: São Paulo, Rio Grande do Sul.

 

obscurus de Saussure

Polistes obscurus de Saussure, 1863, Mém. Soc. Phys. Hist. Nat. Genève 17: 236.

Polistes obscurus var. simulans Bertoni, 1918, An. Cient. Parag. (2) 3: 228.

Distribution: Brazil; Paraguay; Argentina: Misiones (Richards, 1978, Soc. Wasps Am.).

 

occipitalis Ducke

Polistes occipitalis Ducke, 1904, Bol. Mus. Goeldi, Pará 4: 371.

Distribution: Panama; Colombia; Peru; Guyana; Suriname; French Guiana; Brazil: Amapá, Amazonas, Pará, Espírito Santo, Rio de Janeiro, São Paulo; Bolivia (Richards, 1978, Soc. Wasps Am.).

 

oculatus Smith

Polistes oculatus Smith, 1857, Cat. Hym. Br. Mus. 5, Vesp.: 111.

Distribution: Mexico: Guadalajara, Morelos, Oaxaca, Veracruz; Guatemala; Honduras; Costa Rica (Richards, 1978, Soc. Wasps Am.; West-Eberhard et al., 1995, in Hanson and Gauld, Hym. Costa Rica).

 

pacificus Fabricius

Polistes pacifica Fabricius, 1804, Syst. Piez.: 274.

Polistes liliaciosus de Saussure, 1854, Ét. Fam. Vesp. 2: 97.

Polistes liliaceusculus de Saussure, 1854, Ét. Fam. Vesp. 2: 98.

Polistes pacificus flavopictus Ducke, 1918, Rev. Mus. Paulista 10: 360.

Polistes modestus Smith, 1862, Trans. Entomol. Soc. London (3) 1: 38.

Polistes pacificus var. trinitatis Bequaert, 1937, Arch. Inst. Biol. Veg. 3: 197.

Polistes pacificus var. flavopicus [!]; Richards, 1978, Soc. Wasps Am.: 549.

Distribution: U.S.A.: southern Texas; Mexico: Veracruz, Colima, Guerrero, Morelos, Yucatán; Guatemala; Honduras; Nicaragua; Costa Rica; Panama; Colombia; Ecuador; Peru; Venezuela; Trinidad; Guyana; Suriname; French Guiana; Brazil: Amapá to Amazonas, Bahia to São Paulo; Paraguay; Bolivia (Richards, 1978, Soc. Wasps Am.; Snelling, 1983, Pan-Pacif. Entomol. 59).

 

palmarum Bequaert

Polistes major var. palmarum Bequaert, 1936, Entomol. News 47: 11.

Polistes major slevini Bohart, 1949, Pan-Pacif. Entomol. 25: 103.

Distribution: U.S.A.: southern California; Mexico: Baja California (Snelling, 1983, Pan-Pacif. Entomol. 59).

 

paraguayensis Bertoni

Polistes paraguayensis Bertoni, 1921, Rev. Soc. Cient. Paraguay 1: 12.

Distribution: Brazil: Mato Grosso; Bolivia; Paraguay; Argentina: Misiones (Richards, 1978, Soc. Wasps Am.).

 

penai Richards

Polistes penai Richards, 1978, Soc. Wasps Am.: 494.

Distribution: Bolivia.

 

perplexus Cresson

Polistes perplexus Cresson, 1872, Trans. Am. Entomol. Soc. 4: 245.

Polistes generosus Cresson, 1872, Trans. Am. Entomol. Soc. 4: 246.

Distribution: U.S.A.: Maryland to Georgia, west to Kansas, south to Texas (Krombein, 1979, in Krombein et al., Cat. Hym. Am. N. of Mexico).

 

peruvianus Bequaert

Polistes versicolor var. peruvianus Bequaert, 1934, Rev. Entomol., Rio de Janeiro 4: 152.

Distribution: Peru (Richards, 1978, Soc. Wasps Am.).

 

poeyi haitiensis Bequaert and Salt

Polistes poeyi var. haitiensis Bequaert and Salt, 1931, Ann. Entomol. Soc. Am. 24: 792.

Polistes poeyi var. haitensis [!]; Yoshikawa, 1962, J. Biol. Osaka City Univ. 13: 35.

Distribution: Haiti; Dominican Republic.

 

poeyi poeyi Lepeletier

Polistes poeyi Lepeletier, 1836, Hist. Nat. Insect. Hym. 1: 532.

Distribution: Cuba, Is. de la Juventud (Richards, 1978, Soc. Wasps. Am.).

 

pseudoculatus Snelling

Polistes instabilis pseudoculatus Snelling, 1955, Am. Mus. Novitates 1701: 8.

Distribution: Mexico: Chihuahua.

 

ridleyi Kirby

Polistes ridleyi Kirby, 1890, J. Linn. Soc. (Zool.) 20: 541.

Distribution: Brazil: Fernando Noronha.

 

rossi Bohart

Polistes rossi Bohart, 1949, Pan-Pacif. Entomol. 25: 99.

Distribution: Mexico: Baja California.

 

rufidens de Saussure

Polistes rufidens de Saussure, 1853, Ét. Fam. Vesp. 2: 77.

Distribution: Colombia; Venezuela (Richards, 1978, Soc. Wasps Am.).

 

rufiventris Ducke

Polistes rufiventris Ducke, 1904, Bol. Mus. Goeldi, Pará 4: 371.

Polistes erythrogaster Ducke, 1905, Rev. Entomol., Caen 24: 23.

Distribution: Colombia; Guyana; Ecuador; Peru; Brazil: Pará, Amazonas, Rondônia (Richards, 1978, Soc. Wasps Am.).

 

satan Bequaert

Polistes canadensis var. (or subsp.) satan Bequaert, 1940, J. New York Entomol. Soc. 28 : 11.

? Polistes melanarius Zikán, 1951, Dusenia 2: 230.

Distribution: Brazil: Goiás, D. F., Rio de Janeiro, São Paulo, Minas Gerais (Richards, 1978, Soc. Wasps Am.).

 

simillimus Zikán

Polistes simillimus Zikán, 1951, Dusenia 2: 232.

Distribution: Brazil: Bahia to Rio Grande do Sul; Bolivia; Paraguay; Argentina: Jujuy to Misiones, south to Buenos Aires (Richards, 1978, Soc. Wasps Am.).

 

stabilinus Richards

Polistes stabilinus Richards, 1978, Soc. Wasps Am.: 510.

Distribution: Mexico: Sonora, Jalisco, Veracruz, Tabasco, Yucatán; Belize; Honduras.

 

subsericeus de Saussure

Polistes subsericeus de Saussure, 1854, Ét. Fam. Vesp. 2: 100.

Distribution: Colombia; Venezuela; Guyana; Suriname; Brazil: Amapá, Amazonas, Pará to Mato Grosso, São Paulo; Paraguay; Bolivia (Richards, 1978, Soc. Wasps Am.).

 

testaceicolor Bequaert

Vespa analis Fabricius, 1798, Suppl. Entomol. Syst.: 261, non Fabricius, 1775.

Polistes testaceicolor Bequaert, 1937, Arch. Inst. Biol. Veg. 3: 179.  Replacement name for analis Fabricius, 1798, non Fabricius, 1775.

Distribution: Costa Rica; Colombia; Venezuela; Guyana; Suriname; French Guiana; Ecuador; Peru; Brazil: Amapá to Acre, Mato Grosso; Bolivia (Bequaert, 1949, Bol. Mus. Venez. 7; Richards, 1978, Soc. Wasps Am.).

 

thoracicus Fox

Polistes thoracicus Fox, 1898, Proc. Acad. Nat. Sci. Philadelphia 1898: 447.

Distribution: Brazil: Goiás, Matto Grosso; Paraguay (Richards, 1978, Soc. Wasps Am.).

 

torresae Silveira

Polistes occultus Silveira, 1994, Stud. Neotrop. Fauna Env. 29 (4): 189, non Kojima, 1988.

Polistes torresae Silveira, 1996, in Carpenter, Am. Mus. Novitat. 3188: 1, 33.  Replacement name for occultus Silveira, 1994, non Kojima, 1988.

Distribution: Brazil: Pará, Rondônia.

 

veracrucis Richards

Polistes veracrucis Richards, 1978, Soc. Wasps Am.: 508.

Distribution: Mexico: Nuevo León, Veracruz, Hidalgo.

 

versicolor flavoguttatus Bequaert

Polistes versicolor var. flavoguttatus Bequaert, 1934, Rev. Entomol., Rio de Janeiro 4: 155.

Distribution: Ecuador; Peru; Bolivia (Richards, 1978, Soc. Wasps Am.).

 

versicolor kaieteurensis Bequaert

Polistes versicolor var. kaieteurensis Bequaert, 1934, Rev. Entomol., Rio de Janeiro 4: 156.

Distribution: Guyana; Brazil: Roraima (Richards, 1978, Soc. Wasps Am.).

 

versicolor versicolor (Olivier)

Vespa versicolor Olivier, 1792, Encyc. Méthod. 6: 692.

Vespa myops Fabricius, 1798, Suppl. Entomol. Syst.: 261.

Polistes opalinus de Saussure, 1853, Ét. Fam. Vesp. 2: 89.

Polistes versicolor var. vulgaris Bequaert, 1934, Rev. Entomol., Rio de Janeiro 4: 150.

Distribution: Costa Rica; Panama; Colombia; Ecuador; Peru; Venezuela; Trinidad; Grenada; Guyana; Suriname; French Guiana; Brazil: Amapá to Acre, Pará to Rio Grande do Sul, Mato Grosso; Bolivia; Paraguay; Argentina: Jujuy to Misiones, south to Buenos Aires (Richards, 1978, Soc. Wasps Am.).

 

weyrauchorum Willink

Polistes weyrauchorum Willink, 1964, Acta Zool. Lilloana: 12.

Distribution: Ecuador; Peru; Chile (Richards, 1978, Soc. Wasps Am.).

 

xanthogaster willei Bequaert

Polistes versicolor var. (or subsp.) willei Bequaert, 1940, Entomol. News 51: 81.

Distribution: Peru (Richards, 1978, Soc. Wasps Am.).

 

xanthogaster xanthogaster Bequaert

Polistes versicolor var. (or subsp.) xanthogaster Bequaert, 1940, Entomol. News 51: 81.

Distribution: Bolivia; Argentina: Córdoba (Richards, 1978, Soc. Wasps Am.).

 

 

FOSSIL SPECIES

 

attavinus (Heer)

Vespa attavina Heer, 1850, Neue Denkschr. Allg. Schweiz. Ges. Gesammt. Naturwiss. 11: 101, pl. VII fig. 8.

Vespa atavina [!]; Dalla Torre, 1894, Cat. Hym. 9: 137.

? Polistes gprimativa Heerh Piton, 1940, Thesis Fac. Sci., Univ. Clermont: 227, nomen nudum.

Miocene of Austria (Bequaert, 1930, Bull. Brooklyn Entomol. Soc. 25: 69).

industrius Theobald

Polistes industrius Theobald, 1937, Bull. Soc. Sci. Nancy 2: 399, pl. VIII fig. 10, pl. XXVIII fig. 9.

Oligocene of France.

kirbyanus Cockerell

Polistes kirbyanus Cockerell, 1914, Proc. Acad. Nat. Sci. Philadelphia 66: 638.

Miocene of Colorado.

signatus Statz

Polistes signata Statz, 1936, Decheniana 93: 282, pl. XI fig. 33.

Oligocene of Germany.

vergnei Piton

Polistes vergnei Piton, 1940, Thesis Fac. Sci., Univ. Clermont: 226, fig. 93.

Paleocene of France.

 

UNRECOGNIZED TAXA

 

concinnus (Fabricius)

Vespa concinna Fabricius, 1775, Syst. Entomol.: 367.

Distribution: Australia (Schulz, 1906, Spolia Hym.: 231).

fuscus Fabricius

Polistes fusca Fabricius, 1804, Syst. Piez.: 274.

Distribution: Australia.

luctuosus (Smith)

Polybia luctuosa Smith, 1858, J. Proc. Linn. Soc. Zool. 2: 114.

Distribution: Borneo (van der Vecht, 1966, Zool. Verh., Leiden 82).

rubidus Lepeletier

Polistes rubida Lepeletier, 1836, Hist. Nat. Insect. Hym. 1: 523.

Distribution: South Africa.

spinolae de Saussure

Polistes spinolae de Saussure, 1853, Ét. Fam. Vesp. 2: 80.

Distribution: Brazil.

 

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